#METABOLOMICS WORKBENCH Andre_Gollowitzer_20240705_020920 DATATRACK_ID:4981 STUDY_ID:ST003336 ANALYSIS_ID:AN005468 PROJECT_ID:PR002065
VERSION             	1
CREATED_ON             	July 22, 2024, 5:48 pm
#PROJECT
PR:PROJECT_TITLE                 	Changes in phospholipid fatty acid composition under cytotoxic stress facilitate
PR:PROJECT_TITLE                 	peroxidation
PR:PROJECT_SUMMARY               	Programs leading to cell death, such as apoptosis, necroptosis, and ferroptosis,
PR:PROJECT_SUMMARY               	involve an oxidative component linked to lipid metabolism that influences
PR:PROJECT_SUMMARY               	membrane homeostasis. Emerging evidence suggests inter-program cross-talk,
PR:PROJECT_SUMMARY               	emphasizing the need for overarching regulatory mechanisms. We show that under
PR:PROJECT_SUMMARY               	specific cytotoxic stress conditions, exogenous or released polyunsaturated
PR:PROJECT_SUMMARY               	fatty acids (PUFAs) are channeled into overall depleting phospholipids that
PR:PROJECT_SUMMARY               	become vulnerable to peroxidation in the presence of associated redox stress. In
PR:PROJECT_SUMMARY               	fibroblasts, this reprogramming results from reduced growth factor receptor
PR:PROJECT_SUMMARY               	tyrosine kinase (RTK) and phosphatidylinositol-3-kinase (PI3K)/Akt signaling,
PR:PROJECT_SUMMARY               	which reduces de novo fatty acid biosynthesis by mechanisms that differ
PR:PROJECT_SUMMARY               	depending on the specific cytotoxic stressors. We conclude that alterations in
PR:PROJECT_SUMMARY               	PUFA metabolism during cytotoxic stress render cells prone to oxidative
PR:PROJECT_SUMMARY               	modifications in phospholipids.
PR:INSTITUTE                     	University of Innsbruck
PR:DEPARTMENT                    	Michael Popp Institute
PR:LAST_NAME                     	Koeberle
PR:FIRST_NAME                    	Andreas
PR:ADDRESS                       	Mitterweg 24, Innsbruck, Tyrol, 6020, Austria
PR:EMAIL                         	andreas.koeberle@uibk.ac.at
PR:PHONE                         	+43 512 507 57903
PR:FUNDING_SOURCE                	Austrian Science Fund (FWF) (P 36299). German Research Council (GRK 1715 and KO
PR:FUNDING_SOURCE                	4589/4-1), the Phospholipid Research Center Heidelberg (AKO-2015-037/1-1,
PR:FUNDING_SOURCE                	AKO-2019-070/2-1, AKO-2O22-100/2-2), the University of Jena (DRM/2013-05 and
PR:FUNDING_SOURCE                	2.7-05).
PR:CONTRIBUTORS                  	André Gollowitzer, Helmut Pein, Konstantin Loeser, Maria Thuermer, and Andreas
PR:CONTRIBUTORS                  	Koeberle
#STUDY
ST:STUDY_TITLE                   	Differentiation between cytotoxic stress-induced PUFA biosynthesis and PUFA
ST:STUDY_TITLE                   	incorporation into phospholipids by metabolic flux studies in fibroblasts
ST:STUDY_SUMMARY                 	Metabolic flux studies addressing the incorporation of i) deuterium labeled
ST:STUDY_SUMMARY                 	polyunsaturated fatty acids (20:4, 22:4, 22:5, 22:6) having one or two
ST:STUDY_SUMMARY                 	acetate-d3 incorporated or ii) one arachidonic acid-d8 or its elongation product
ST:STUDY_SUMMARY                 	adrenic acid-d8 into cellular phosphatidylcholines of NIH-3T3 fibroblasts that
ST:STUDY_SUMMARY                 	were treated with TOFA, VAL or a combination of both
ST:INSTITUTE                     	University of Innsbruck
ST:DEPARTMENT                    	Michael Popp Institute
ST:LAST_NAME                     	Koeberle
ST:FIRST_NAME                    	Andreas
ST:ADDRESS                       	Mitterweg 24, Innsbruck, Tyrol, 6020, Austria
ST:EMAIL                         	andreas.koeberle@uibk.ac.at
ST:PHONE                         	+43 512 507 57903
#SUBJECT
SU:SUBJECT_TYPE                  	Cultured cells
SU:SUBJECT_SPECIES               	Mus musculus
SU:TAXONOMY_ID                   	10090
#SUBJECT_SAMPLE_FACTORS:         	SUBJECT(optional)[tab]SAMPLE[tab]FACTORS(NAME:VALUE pairs separated by |)[tab]Raw file names and additional sample data
SUBJECT_SAMPLE_FACTORS           	-	200705_PUFA_PC_dPUFA_dAA_n_1_DMSO_1	Sample source:cultured cells | treatment A:vehicle (DMSO) | treatment B:- | treatment C:-	RAW_FILE_NAME(Local_data_file)=200705_PUFA_PC_dPUFA_dAA_n_1; concentration A=-; concentration B=-; concentration C=-
SUBJECT_SAMPLE_FACTORS           	-	200705_PUFA_PC_dPUFA_dAA_n_1_DMSO_dAA_2	Sample source:cultured cells | treatment A:vehicle (DMSO) | treatment B:- | treatment C:dAA	RAW_FILE_NAME(Local_data_file)=200705_PUFA_PC_dPUFA_dAA_n_1; concentration A=-; concentration B=-; concentration C=1 µM
SUBJECT_SAMPLE_FACTORS           	-	200705_PUFA_PC_dPUFA_dAA_n_1_TOFA_dAA_4	Sample source:cultured cells | treatment A:TOFA | treatment B:- | treatment C:dAA	RAW_FILE_NAME(Local_data_file)=200705_PUFA_PC_dPUFA_dAA_n_1; concentration A=5 µM; concentration B=-; concentration C=1 µM
SUBJECT_SAMPLE_FACTORS           	-	200705_PUFA_PC_dPUFA_dAA_n_1_VAL_dAA_6	Sample source:cultured cells | treatment A:VAL | treatment B:- | treatment C:dAA	RAW_FILE_NAME(Local_data_file)=200705_PUFA_PC_dPUFA_dAA_n_1; concentration A=10 µM; concentration B=-; concentration C=1 µM
SUBJECT_SAMPLE_FACTORS           	-	200705_PUFA_PC_dPUFA_dAA_n_1_TOFA_VAL_dAA_8	Sample source:cultured cells | treatment A:TOFA | treatment B:VAL | treatment C:dAA	RAW_FILE_NAME(Local_data_file)=200705_PUFA_PC_dPUFA_dAA_n_1; concentration A=5 µM; concentration B=10 µM; concentration C=1 µM
SUBJECT_SAMPLE_FACTORS           	-	200709_PUFA_PC_dPUFA_dAa_n2_DMSO	Sample source:cultured cells | treatment A:vehicle (DMSO) | treatment B:- | treatment C:-	RAW_FILE_NAME(Local_data_file)=200709_PUFA_PC_dPUFA_dAa_n2_n3_dil_1_2; concentration A=-; concentration B=-; concentration C=-
SUBJECT_SAMPLE_FACTORS           	-	200709_PUFA_PC_dPUFA_dAa_n2_DMSO_dAa	Sample source:cultured cells | treatment A:vehicle (DMSO) | treatment B:- | treatment C:dAA	RAW_FILE_NAME(Local_data_file)=200709_PUFA_PC_dPUFA_dAa_n2_n3_dil_1_2; concentration A=-; concentration B=-; concentration C=1 µM
SUBJECT_SAMPLE_FACTORS           	-	200709_PUFA_PC_dPUFA_dAa_n2_TOFA_dAa	Sample source:cultured cells | treatment A:TOFA | treatment B:- | treatment C:dAA	RAW_FILE_NAME(Local_data_file)=200709_PUFA_PC_dPUFA_dAa_n2_n3_dil_1_2; concentration A=5 µM; concentration B=-; concentration C=1 µM
SUBJECT_SAMPLE_FACTORS           	-	200709_PUFA_PC_dPUFA_dAa_n2_VAL_dAa	Sample source:cultured cells | treatment A:VAL | treatment B:- | treatment C:dAA	RAW_FILE_NAME(Local_data_file)=200709_PUFA_PC_dPUFA_dAa_n2_n3_dil_1_2; concentration A=10 µM; concentration B=-; concentration C=1 µM
SUBJECT_SAMPLE_FACTORS           	-	200709_PUFA_PC_dPUFA_dAa_n2_TOFA_VAL_dAa	Sample source:cultured cells | treatment A:TOFA | treatment B:VAL | treatment C:dAA	RAW_FILE_NAME(Local_data_file)=200709_PUFA_PC_dPUFA_dAa_n2_n3_dil_1_2; concentration A=5 µM; concentration B=10 µM; concentration C=1 µM
SUBJECT_SAMPLE_FACTORS           	-	200709_PUFA_PC_dPUFA_dAa_n3_DMSO	Sample source:cultured cells | treatment A:vehicle (DMSO) | treatment B:- | treatment C:-	RAW_FILE_NAME(Local_data_file)=200709_PUFA_PC_dPUFA_dAa_n2_n3_dil_1_2; concentration A=-; concentration B=-; concentration C=-
SUBJECT_SAMPLE_FACTORS           	-	200709_PUFA_PC_dPUFA_dAa_n3_DMSO_dAa	Sample source:cultured cells | treatment A:vehicle (DMSO) | treatment B:- | treatment C:dAA	RAW_FILE_NAME(Local_data_file)=200709_PUFA_PC_dPUFA_dAa_n2_n3_dil_1_2; concentration A=-; concentration B=-; concentration C=1 µM
SUBJECT_SAMPLE_FACTORS           	-	200709_PUFA_PC_dPUFA_dAa_n3_TOFA_dAa	Sample source:cultured cells | treatment A:TOFA | treatment B:- | treatment C:dAA	RAW_FILE_NAME(Local_data_file)=200709_PUFA_PC_dPUFA_dAa_n2_n3_dil_1_2; concentration A=5 µM; concentration B=-; concentration C=1 µM
SUBJECT_SAMPLE_FACTORS           	-	200709_PUFA_PC_dPUFA_dAa_n3_VAL_dAa	Sample source:cultured cells | treatment A:VAL | treatment B:- | treatment C:dAA	RAW_FILE_NAME(Local_data_file)=200709_PUFA_PC_dPUFA_dAa_n2_n3_dil_1_2; concentration A=10 µM; concentration B=-; concentration C=1 µM
SUBJECT_SAMPLE_FACTORS           	-	200709_PUFA_PC_dPUFA_dAa_n3_TOFA_VAL_dAa	Sample source:cultured cells | treatment A:TOFA | treatment B:VAL | treatment C:dAA	RAW_FILE_NAME(Local_data_file)=200709_PUFA_PC_dPUFA_dAa_n2_n3_dil_1_2; concentration A=5 µM; concentration B=10 µM; concentration C=1 µM
SUBJECT_SAMPLE_FACTORS           	-	200705_PUFA_PC_dPUFA_dAC_n_1_DMSO_1	Sample source:cultured cells | treatment A:vehicle (DMSO) | treatment B:- | treatment C:-	RAW_FILE_NAME(Local_data_file)=200705_PUFA_PC_dPUFA_dAC_n_1; concentration A=-; concentration B=-; concentration C=-
SUBJECT_SAMPLE_FACTORS           	-	200705_PUFA_PC_dPUFA_dAC_n_1_DMSO_dAc_2	Sample source:cultured cells | treatment A:vehicle (DMSO) | treatment B:- | treatment C:dAc	RAW_FILE_NAME(Local_data_file)=200705_PUFA_PC_dPUFA_dAC_n_1; concentration A=-; concentration B=-; concentration C=30 µM
SUBJECT_SAMPLE_FACTORS           	-	200705_PUFA_PC_dPUFA_dAC_n_1_TOFA_dAc_4	Sample source:cultured cells | treatment A:TOFA | treatment B:- | treatment C:dAc	RAW_FILE_NAME(Local_data_file)=200705_PUFA_PC_dPUFA_dAC_n_1; concentration A=5 µM; concentration B=-; concentration C=30 µM
SUBJECT_SAMPLE_FACTORS           	-	200705_PUFA_PC_dPUFA_dAC_n_1_VAL_dAc_6	Sample source:cultured cells | treatment A:VAL | treatment B:- | treatment C:dAc	RAW_FILE_NAME(Local_data_file)=200705_PUFA_PC_dPUFA_dAC_n_1; concentration A=10 µM; concentration B=-; concentration C=30 µM
SUBJECT_SAMPLE_FACTORS           	-	200705_PUFA_PC_dPUFA_dAC_n_1_TOFA_VAL_dAc_8	Sample source:cultured cells | treatment A:TOFA | treatment B:VAL | treatment C:dAc	RAW_FILE_NAME(Local_data_file)=200705_PUFA_PC_dPUFA_dAC_n_1; concentration A=5 µM; concentration B=10 µM; concentration C=30 µM
SUBJECT_SAMPLE_FACTORS           	-	200709_PUFA_PC_dPUFA_dAc_n2_DMSO	Sample source:cultured cells | treatment A:vehicle (DMSO) | treatment B:- | treatment C:-	RAW_FILE_NAME(Local_data_file)=200716_PUFA_PC_dPUFA_dAc_n2_n3_dil_1_2; concentration A=-; concentration B=-; concentration C=-
SUBJECT_SAMPLE_FACTORS           	-	200709_PUFA_PC_dPUFA_dAc_n2_DMSO_dAc	Sample source:cultured cells | treatment A:vehicle (DMSO) | treatment B:- | treatment C:dAc	RAW_FILE_NAME(Local_data_file)=200716_PUFA_PC_dPUFA_dAc_n2_n3_dil_1_2; concentration A=-; concentration B=-; concentration C=30 µM
SUBJECT_SAMPLE_FACTORS           	-	200709_PUFA_PC_dPUFA_dAc_n2_TOFA_dAc	Sample source:cultured cells | treatment A:TOFA | treatment B:- | treatment C:dAc	RAW_FILE_NAME(Local_data_file)=200716_PUFA_PC_dPUFA_dAc_n2_n3_dil_1_2; concentration A=5 µM; concentration B=-; concentration C=30 µM
SUBJECT_SAMPLE_FACTORS           	-	200709_PUFA_PC_dPUFA_dAc_n2_VAL_dAc	Sample source:cultured cells | treatment A:VAL | treatment B:- | treatment C:dAc	RAW_FILE_NAME(Local_data_file)=200716_PUFA_PC_dPUFA_dAc_n2_n3_dil_1_2; concentration A=10 µM; concentration B=-; concentration C=30 µM
SUBJECT_SAMPLE_FACTORS           	-	200709_PUFA_PC_dPUFA_dAc_n2_TOFA_VAL_dAc	Sample source:cultured cells | treatment A:TOFA | treatment B:VAL | treatment C:dAc	RAW_FILE_NAME(Local_data_file)=200716_PUFA_PC_dPUFA_dAc_n2_n3_dil_1_2; concentration A=5 µM; concentration B=10 µM; concentration C=30 µM
SUBJECT_SAMPLE_FACTORS           	-	200709_PUFA_PC_dPUFA_dAc_n3_DMSO	Sample source:cultured cells | treatment A:vehicle (DMSO) | treatment B:- | treatment C:-	RAW_FILE_NAME(Local_data_file)=200716_PUFA_PC_dPUFA_dAc_n2_n3_dil_1_2; concentration A=-; concentration B=-; concentration C=-
SUBJECT_SAMPLE_FACTORS           	-	200709_PUFA_PC_dPUFA_dAc_n3_DMSO_dAc	Sample source:cultured cells | treatment A:vehicle (DMSO) | treatment B:- | treatment C:dAc	RAW_FILE_NAME(Local_data_file)=200716_PUFA_PC_dPUFA_dAc_n2_n3_dil_1_2; concentration A=-; concentration B=-; concentration C=30 µM
SUBJECT_SAMPLE_FACTORS           	-	200709_PUFA_PC_dPUFA_dAc_n3_TOFA_dAc	Sample source:cultured cells | treatment A:TOFA | treatment B:- | treatment C:dAc	RAW_FILE_NAME(Local_data_file)=200716_PUFA_PC_dPUFA_dAc_n2_n3_dil_1_2; concentration A=5 µM; concentration B=-; concentration C=30 µM
SUBJECT_SAMPLE_FACTORS           	-	200709_PUFA_PC_dPUFA_dAc_n3_VAL_dAc	Sample source:cultured cells | treatment A:VAL | treatment B:- | treatment C:dAc	RAW_FILE_NAME(Local_data_file)=200716_PUFA_PC_dPUFA_dAc_n2_n3_dil_1_2; concentration A=10 µM; concentration B=-; concentration C=30 µM
SUBJECT_SAMPLE_FACTORS           	-	200709_PUFA_PC_dPUFA_dAc_n3_TOFA_VAL_dAc	Sample source:cultured cells | treatment A:TOFA | treatment B:VAL | treatment C:dAc	RAW_FILE_NAME(Local_data_file)=200716_PUFA_PC_dPUFA_dAc_n2_n3_dil_1_2; concentration A=5 µM; concentration B=10 µM; concentration C=30 µM
#COLLECTION
CO:COLLECTION_SUMMARY            	Cultured cells were washed, trypsinized, counted and flash-frozen in liquid N2
CO:COLLECTION_SUMMARY            	and stored at -80°C.
CO:SAMPLE_TYPE                   	Fibroblasts
CO:COLLECTION_METHOD             	Trypsinization of cultured cells
CO:STORAGE_CONDITIONS            	-80℃
#TREATMENT
TR:TREATMENT_SUMMARY             	For metabolic flux studies, NIH-3T3 cells were incubated with acetate-d3 (30
TR:TREATMENT_SUMMARY             	µM) or arachidonic acid-d8 (1 µM) and treated with vehicle (DMSO), TOFA (5
TR:TREATMENT_SUMMARY             	µM), VAL (10 µM), or both VAL and TOFA.
#SAMPLEPREP
SP:SAMPLEPREP_SUMMARY            	Phospholipids were extracted from cell pellets by successive addition of PBS pH
SP:SAMPLEPREP_SUMMARY            	7.4, methanol, chloroform, and saline to a final ratio of 14:34:35:17.
SP:SAMPLEPREP_SUMMARY            	Evaporation of the organic layer yielded a lipid film that was dissolved in
SP:SAMPLEPREP_SUMMARY            	methanol, diluted, and subjected to UPLC-MS/MS.
SP:EXTRACT_STORAGE               	-20℃
#CHROMATOGRAPHY
CH:CHROMATOGRAPHY_SUMMARY        	Chromatographic separation of (deuterated) phospholipids was carried out on an
CH:CHROMATOGRAPHY_SUMMARY        	Acquity BEH C8 column (1.7 μm, 2.1×100 mm, Waters, Milford, MA) using an
CH:CHROMATOGRAPHY_SUMMARY        	ExionLC AD UHPLC system (Sciex).
CH:CHROMATOGRAPHY_TYPE           	Reversed phase
CH:INSTRUMENT_NAME               	Sciex ExionLC AD
CH:COLUMN_NAME                   	Waters ACQUITY UPLC BEH C8 (100 x 2.1mm,1.7um)
CH:SOLVENT_A                     	5% Water/95% Acetonitrile; 2 mM ammonium acetate
CH:SOLVENT_B                     	90% Water/10% Acetonitrile; 2 mM ammonium acetate
CH:FLOW_GRADIENT                 	The gradient was ramped from 75 to 85% A over 5 min and further increased to
CH:FLOW_GRADIENT                 	100% A within 2 min, followed by isocratic elution for another 2 min.
CH:FLOW_RATE                     	0.75 ml/min
CH:COLUMN_TEMPERATURE            	45
#ANALYSIS
AN:ANALYSIS_TYPE                 	MS
#MS
MS:INSTRUMENT_NAME               	ABI Sciex 6500+
MS:INSTRUMENT_TYPE               	QTRAP
MS:MS_TYPE                       	ESI
MS:ION_MODE                      	NEGATIVE
MS:MS_COMMENTS                   	Targeted MRM with pre-optimized settings and subsequent automated integration of
MS:MS_COMMENTS                   	selected signals using Analyst 1.6 and 1.7 (Sciex).
#MS_METABOLITE_DATA
MS_METABOLITE_DATA:UNITS	nmol / 10^6 cells
MS_METABOLITE_DATA_START
Samples	200705_PUFA_PC_dPUFA_dAA_n_1_DMSO_1	200705_PUFA_PC_dPUFA_dAA_n_1_DMSO_dAA_2	200705_PUFA_PC_dPUFA_dAA_n_1_TOFA_dAA_4	200705_PUFA_PC_dPUFA_dAA_n_1_VAL_dAA_6	200705_PUFA_PC_dPUFA_dAA_n_1_TOFA_VAL_dAA_8	200709_PUFA_PC_dPUFA_dAa_n2_DMSO	200709_PUFA_PC_dPUFA_dAa_n2_DMSO_dAa	200709_PUFA_PC_dPUFA_dAa_n2_TOFA_dAa	200709_PUFA_PC_dPUFA_dAa_n2_VAL_dAa	200709_PUFA_PC_dPUFA_dAa_n2_TOFA_VAL_dAa	200709_PUFA_PC_dPUFA_dAa_n3_DMSO	200709_PUFA_PC_dPUFA_dAa_n3_DMSO_dAa	200709_PUFA_PC_dPUFA_dAa_n3_TOFA_dAa	200709_PUFA_PC_dPUFA_dAa_n3_VAL_dAa	200709_PUFA_PC_dPUFA_dAa_n3_TOFA_VAL_dAa	200705_PUFA_PC_dPUFA_dAC_n_1_DMSO_1	200705_PUFA_PC_dPUFA_dAC_n_1_DMSO_dAc_2	200705_PUFA_PC_dPUFA_dAC_n_1_TOFA_dAc_4	200705_PUFA_PC_dPUFA_dAC_n_1_VAL_dAc_6	200705_PUFA_PC_dPUFA_dAC_n_1_TOFA_VAL_dAc_8	200709_PUFA_PC_dPUFA_dAc_n2_DMSO	200709_PUFA_PC_dPUFA_dAc_n2_DMSO_dAc	200709_PUFA_PC_dPUFA_dAc_n2_TOFA_dAc	200709_PUFA_PC_dPUFA_dAc_n2_VAL_dAc	200709_PUFA_PC_dPUFA_dAc_n2_TOFA_VAL_dAc	200709_PUFA_PC_dPUFA_dAc_n3_DMSO	200709_PUFA_PC_dPUFA_dAc_n3_DMSO_dAc	200709_PUFA_PC_dPUFA_dAc_n3_TOFA_dAc	200709_PUFA_PC_dPUFA_dAc_n3_VAL_dAc	200709_PUFA_PC_dPUFA_dAc_n3_TOFA_VAL_dAc
Factors	Sample source:cultured cells | treatment A:vehicle (DMSO) | treatment B:- | treatment C:-	Sample source:cultured cells | treatment A:vehicle (DMSO) | treatment B:- | treatment C:dAA	Sample source:cultured cells | treatment A:TOFA | treatment B:- | treatment C:dAA	Sample source:cultured cells | treatment A:VAL | treatment B:- | treatment C:dAA	Sample source:cultured cells | treatment A:TOFA | treatment B:VAL | treatment C:dAA	Sample source:cultured cells | treatment A:vehicle (DMSO) | treatment B:- | treatment C:-	Sample source:cultured cells | treatment A:vehicle (DMSO) | treatment B:- | treatment C:dAA	Sample source:cultured cells | treatment A:TOFA | treatment B:- | treatment C:dAA	Sample source:cultured cells | treatment A:VAL | treatment B:- | treatment C:dAA	Sample source:cultured cells | treatment A:TOFA | treatment B:VAL | treatment C:dAA	Sample source:cultured cells | treatment A:vehicle (DMSO) | treatment B:- | treatment C:-	Sample source:cultured cells | treatment A:vehicle (DMSO) | treatment B:- | treatment C:dAA	Sample source:cultured cells | treatment A:TOFA | treatment B:- | treatment C:dAA	Sample source:cultured cells | treatment A:VAL | treatment B:- | treatment C:dAA	Sample source:cultured cells | treatment A:TOFA | treatment B:VAL | treatment C:dAA	Sample source:cultured cells | treatment A:vehicle (DMSO) | treatment B:- | treatment C:-	Sample source:cultured cells | treatment A:vehicle (DMSO) | treatment B:- | treatment C:dAc	Sample source:cultured cells | treatment A:TOFA | treatment B:- | treatment C:dAc	Sample source:cultured cells | treatment A:VAL | treatment B:- | treatment C:dAc	Sample source:cultured cells | treatment A:TOFA | treatment B:VAL | treatment C:dAc	Sample source:cultured cells | treatment A:vehicle (DMSO) | treatment B:- | treatment C:-	Sample source:cultured cells | treatment A:vehicle (DMSO) | treatment B:- | treatment C:dAc	Sample source:cultured cells | treatment A:TOFA | treatment B:- | treatment C:dAc	Sample source:cultured cells | treatment A:VAL | treatment B:- | treatment C:dAc	Sample source:cultured cells | treatment A:TOFA | treatment B:VAL | treatment C:dAc	Sample source:cultured cells | treatment A:vehicle (DMSO) | treatment B:- | treatment C:-	Sample source:cultured cells | treatment A:vehicle (DMSO) | treatment B:- | treatment C:dAc	Sample source:cultured cells | treatment A:TOFA | treatment B:- | treatment C:dAc	Sample source:cultured cells | treatment A:VAL | treatment B:- | treatment C:dAc	Sample source:cultured cells | treatment A:TOFA | treatment B:VAL | treatment C:dAc
PC(14:0_16:0)	0.06461	0.06711	0.07100	0.19679	0.15427	0.12162	0.10893	0.10568	0.30183	0.17351	0.08918	0.10794	0.09306	0.22486	0.15305															
PC(16:0/16:0)	0.09808	0.10383	0.07173	0.28800	0.35706	0.22280	0.18485	0.10374	0.47953	0.43459	0.16514	0.19971	0.09798	0.36304	0.41561															
PC(16:0_16:1)	0.21185	0.22356	0.09500	0.29903	0.35853	0.39182	0.31362	0.09672	0.47410	0.45619	0.29563	0.31878	0.09600	0.37709	0.38125															
PC(16:1/16:1)	0.08474	0.08162	0.01665	0.06244	0.06042	0.12981	0.10619	0.02764	0.10482	0.08181	0.09788	0.10226	0.02539	0.08500	0.06829															
PC(16:0_18:1)	0.12816	0.12651	0.08658	0.22423	0.29705	0.26949	0.22145	0.12233	0.50923	0.48755	0.19499	0.22911	0.11916	0.41534	0.43717					f										
PC(16:1_18:1)	0.14532	0.14760	0.04601	0.18577	0.21852	0.24803	0.18542	0.06469	0.27036	0.24626	0.19300	0.19880	0.06367	0.23157	0.20490															
PC(18:0_18:1)	0.13006	0.12537	0.08385	0.22420	0.24987	0.24049	0.20463	0.10500	0.50345	0.42783	0.18679	0.21648	0.10213	0.39750	0.35199															
PC(18:1/18:1)	0.18240	0.17480	0.08534	0.16488	0.23279	0.34410	0.28609	0.16749	0.37954	0.37510	0.25834	0.30075	0.15644	0.32208	0.31135															
PC(18:1_18:2)	0.02521	0.02426	0.01623	0.03463	0.04914	0.04497	0.03328	0.03363	0.05980	0.08065	0.03353	0.03534	0.03229	0.05181	0.06499															
PC(16:0_20:4)	0.00551	0.00705	0.01717	0.03770	0.09956	0.02357	0.01851	0.03299	0.06573	0.15206	0.01490	0.02029	0.02833	0.05443	0.13372															
PC(16:0_20:4-d8)	0.00000	0.00084	0.00289	0.00729	0.02500	0.00000	0.00211	0.00522	0.01345	0.03006	0.00001	0.00238	0.00520	0.01138	0.02980															
PC(16:1_20:4)	0.00292	0.00361	0.00329	0.00883	0.01914	0.00957	0.00869	0.01211	0.01421	0.02876	0.00614	0.00808	0.00948	0.01230	0.02439															
PC(16:1_20:4-d8)	0.00000	0.00034	0.00051	0.00185	0.00473	0.00000	0.00081	0.00180	0.00270	0.00507	0.00000	0.00082	0.00153	0.00224	0.00504															
PC(18:0_20:4)	0.00227	0.00277	0.00698	0.01902	0.04585	0.01188	0.01050	0.01908	0.04386	0.09187	0.00745	0.01032	0.01523	0.03303	0.08184															
PC(18:0_20:4-d8)	0.00000	0.00023	0.00085	0.00357	0.01075	0.00002	0.00093	0.00225	0.00670	0.01403	0.00001	0.00095	0.00207	0.00528	0.01515															
PC(18:1_20:4)	0.00644	0.00832	0.01096	0.03397	0.07714	0.02477	0.01827	0.03427	0.04427	0.09868	0.01545	0.01975	0.02909	0.04015	0.08511															
PC(18:1_20:4-d8)	0.00000	0.00070	0.00170	0.00643	0.01757	0.00000	0.00188	0.00493	0.00835	0.01768	0.00001	0.00207	0.00458	0.00664	0.01822															
PC(16:0_22:4)	0.00039	0.00054	0.00088	0.00231	0.00427	0.00222	0.00209	0.00273	0.00848	0.01301	0.00142	0.00213	0.00237	0.00602	0.01092															
PC(16:0_22:4-d8)	0.00000	0.00005	0.00002	0.00014	0.00001	0.00000	0.00026	0.00008	0.00026	0.00001	0.00000	0.00024	0.00005	0.00017	0.00002															
PC(18:0_22:4)	0.00094	0.00121	0.00139	0.00676	0.01103	0.00538	0.00503	0.00387	0.02331	0.03816	0.00353	0.00489	0.00334	0.01864	0.02650															
PC(18:0_22:4-d8)	0.00000	0.00015	0.00002	0.00034	0.00005	0.00000	0.00058	0.00009	0.00055	0.00007	0.00000	0.00046	0.00004	0.00035	0.00001															
PC(18:1_22:4)	0.00040	0.00056	0.00061	0.00177	0.00291	0.00238	0.00227	0.00299	0.00615	0.00903	0.00150	0.00225	0.00223	0.00480	0.00746															
PC(18:1_22:4-d8)	0.00000	0.00005	0.00001	0.00004	0.00001	0.00000	0.00021	0.00005	0.00011	0.00000	0.00000	0.00024	0.00004	0.00012	0.00002															
PC(16:0_22:5)	0.00222	0.00288	0.00529	0.01264	0.02971	0.00733	0.00550	0.00817	0.02593	0.04645	0.00467	0.00565	0.00734	0.01892	0.03290															
PC(18:0_22:5)	0.00068	0.00085	0.00152	0.00633	0.01223	0.00279	0.00238	0.00362	0.01479	0.02429	0.00187	0.00247	0.00278	0.01121	0.01992															
PC(18:1_22:5)	0.00154	0.00173	0.00222	0.00533	0.01047	0.00473	0.00344	0.00560	0.00948	0.01588	0.00316	0.00369	0.00467	0.00808	0.01210															
PC(16:0_22:6)	0.00233	0.00324	0.00529	0.00715	0.01528	0.00784	0.00611	0.00791	0.01458	0.02182	0.00509	0.00628	0.00634	0.01018	0.01749															
PC(16:1_22:6)	0.00076	0.00083	0.00080	0.00150	0.00239	0.00197	0.00156	0.00212	0.00284	0.00378	0.00118	0.00162	0.00163	0.00231	0.00284															
PC(18:0_22:6)	0.00040	0.00049	0.00119	0.00223	0.00410	0.00158	0.00139	0.00219	0.00457	0.00741	0.00094	0.00147	0.00189	0.00356	0.00534															
PC(18:1_22:6)	0.00146	0.00200	0.00297	0.00431	0.00853	0.00538	0.00374	0.00644	0.00796	0.01084	0.00335	0.00398	0.00537	0.00660	0.00881															
PC(16:0_20:4-d1)																0.00000	0.00000	0.00004	0.00000	0.00000	0.00000	0.00001	0.00000	0.00000	0.00000	0.00000	0.00002	0.00000	0.00067	0.00096
PC(16:1_20:4-d1)																0.00000	0.00000	0.00002	0.00000	0.00000	0.00000	0.00000	0.00000	0.00000	0.00000	0.00000	0.00000	0.00000	0.00000	0.00036
PC(18:0_20:4-d1)																0.00000	0.00002	0.00011	0.00000	0.00010	0.00000	0.00006	0.00000	0.00000	0.00000	0.00000	0.00000	0.00072	0.00000	0.00189
PC(18:1_20:4-d1)																0.00000	0.00000	0.00000	0.00000	0.00008	0.00000	0.00002	0.00000	0.00000	0.00000	0.00000	0.00000	0.00000	0.00000	0.00000
PC(16:0_22:4-d1)																0.00000	0.00002	0.00002	0.00000	0.00000	0.00000	0.00000	0.00000	0.00000	0.00000	0.00000	0.00000	0.00000	0.00000	0.00000
PC(16:0_22:4-d2)																0.00000	0.00001	0.00000	0.00002	0.00000	0.00000	0.00001	0.00005	0.00000	0.00001	0.00000	0.00000	0.00000	0.00000	0.00007
PC(18:0_22:4-d1)																0.00000	0.00001	0.00000	0.00007	0.00000	0.00000	0.00000	0.00000	0.00000	0.00000	0.00000	0.00000	0.00000	0.00000	0.00000
PC(18:0_22:4-d2)																0.00000	0.00001	0.00003	0.00003	0.00018	0.00000	0.00003	0.00000	0.00000	0.00000	0.00000	0.00003	0.00004	0.00000	0.00019
PC(18:1_22:4-d1)																0.00000	0.00000	0.00003	0.00006	0.00000	0.00000	0.00000	0.00000	0.00000	0.00000	0.00000	0.00000	0.00000	0.00000	0.00041
PC(18:1_22:4-d2)																0.00000	0.00000	0.00000	0.00002	0.00000	0.00000	0.00000	0.00003	0.00000	0.00000	0.00000	0.00000	0.00000	0.00000	0.00001
PC(16:0_22:5-d1)																0.00000	0.00000	0.00000	0.00000	0.00000	0.00000	0.00012	0.00008	0.00000	0.00000	0.00000	0.00000	0.00000	0.00000	0.00000
PC(16:0_22:5-d2)																0.00000	0.00000	0.00002	0.00014	0.00000	0.00000	0.00001	0.00000	0.00000	0.00011	0.00000	0.00000	0.00000	0.00014	0.00006
PC(18:0_22:5-d1)																0.00000	0.00003	0.00000	0.00000	0.00000	0.00000	0.00000	0.00000	0.00000	0.00000	0.00000	0.00000	0.00000	0.00012	0.00000
PC(18:0_22:5-d2)																0.00000	0.00003	0.00007	0.00010	0.00027	0.00000	0.00006	0.00006	0.00020	0.00035	0.00000	0.00011	0.00019	0.00016	0.00072
PC(18:1_22:5-d1)																0.00000	0.00001	0.00000	0.00000	0.00000	0.00000	0.00001	0.00000	0.00011	0.00000	0.00000	0.00000	0.00000	0.00006	0.00000
PC(18:1_22:5-d2)																0.00000	0.00002	0.00001	0.00002	0.00000	0.00000	0.00002	0.00000	0.00003	0.00001	0.00000	0.00000	0.00003	0.00000	0.00004
PC(16:0_22:6-d1)																0.00000	0.00000	0.00000	0.00000	0.00000	0.00000	0.00008	0.00000	0.00000	0.00000	0.00000	0.00000	0.00000	0.00000	0.00070
PC(16:0_22:6-d2)																0.00000	0.00001	0.00000	0.00000	0.00000	0.00000	0.00000	0.00003	0.00006	0.00013	0.00000	0.00000	0.00000	0.00000	0.00011
PC(18:0_22:6-d1)																0.00000	0.00002	0.00003	0.00000	0.00000	0.00000	0.00001	0.00000	0.00000	0.00035	0.00000	0.00000	0.00000	0.00006	0.00000
PC(18:0_22:6-d2)																0.00000	0.00000	0.00000	0.00000	0.00006	0.00000	0.00001	0.00003	0.00006	0.00004	0.00000	0.00002	0.00000	0.00004	0.00008
PC(18:1_22:6-d1)																0.00000	0.00000	0.00000	0.00009	0.00002	0.00000	0.00002	0.00000	0.00018	0.00044	0.00000	0.00006	0.00000	0.00000	0.00000
PC(18:1_22:6-d2)																0.00000	0.00001	0.00000	0.00000	0.00003	0.00000	0.00000	0.00000	0.00012	0.00004	0.00000	0.00000	0.00000	0.00000	0.00000
MS_METABOLITE_DATA_END
#METABOLITES
METABOLITES_START
metabolite_name	Standardized name	Exact mass	Formula	Super class	Main class	Sub class
PC(14:0_16:0)	PC 14:0_16:0	705.5309	C38H76NO8P	Glycerophospholipids	Glycerophosphocholines	PC
PC(16:0/16:0)	PC 16:0/16:0	733.5622	C40H80NO8P	Glycerophospholipids	Glycerophosphocholines	PC
PC(16:0_16:1)	PC 16:0_16:1	731.5465	C40H78NO8P	Glycerophospholipids	Glycerophosphocholines	PC
PC(16:1/16:1)	PC 16:1/16:1	729.5309	C40H76NO8P	Glycerophospholipids	Glycerophosphocholines	PC
PC(16:0_18:1)	PC 16:0_18:1	759.5778	C42H82NO8P	Glycerophospholipids	Glycerophosphocholines	PC
PC(16:1_18:1)	PC 16:1_18:1	757.5622	C42H80NO8P	Glycerophospholipids	Glycerophosphocholines	PC
PC(18:0_18:1)	PC 18:0_18:1	787.6091	C44H86NO8P	Glycerophospholipids	Glycerophosphocholines	PC
PC(18:1/18:1)	PC 18:1/18:1	785.5935	C44H84NO8P	Glycerophospholipids	Glycerophosphocholines	PC
PC(18:1_18:2)	PC 18:1_18:2	783.5778	C44H82NO8P	Glycerophospholipids	Glycerophosphocholines	PC
PC(16:0_20:4)	PC 16:0_20:4	781.5622	C44H80NO8P	Glycerophospholipids	Glycerophosphocholines	PC
PC(16:0_20:4-d8)	-			Glycerophospholipids	Glycerophosphocholines	PC
PC(16:1_20:4)	PC 16:1_20:4	779.5465	C44H78NO8P	Glycerophospholipids	Glycerophosphocholines	PC
PC(16:1_20:4-d8)	-			Glycerophospholipids	Glycerophosphocholines	PC
PC(18:0_20:4)	PC 18:0_20:4	809.5935	C46H84NO8P	Glycerophospholipids	Glycerophosphocholines	PC
PC(18:0_20:4-d8)	-			Glycerophospholipids	Glycerophosphocholines	PC
PC(18:1_20:4)	PC 18:1_20:4	807.5778	C46H82NO8P	Glycerophospholipids	Glycerophosphocholines	PC
PC(18:1_20:4-d8)	-			Glycerophospholipids	Glycerophosphocholines	PC
PC(16:0_22:4)	PC 16:0_22:4	809.5935	C46H84NO8P	Glycerophospholipids	Glycerophosphocholines	PC
PC(16:0_22:4-d8)	-			Glycerophospholipids	Glycerophosphocholines	PC
PC(18:0_22:4)	PC 18:0_22:4	837.6248	C48H88NO8P	Glycerophospholipids	Glycerophosphocholines	PC
PC(18:0_22:4-d8)	-			Glycerophospholipids	Glycerophosphocholines	PC
PC(18:1_22:4)	PC 18:1_22:4	835.6091	C48H86NO8P	Glycerophospholipids	Glycerophosphocholines	PC
PC(18:1_22:4-d8)	-			Glycerophospholipids	Glycerophosphocholines	PC
PC(16:0_22:5)	PC 16:0_22:5	807.5778	C46H82NO8P	Glycerophospholipids	Glycerophosphocholines	PC
PC(18:0_22:5)	PC 18:0_22:5	835.6091	C48H86NO8P	Glycerophospholipids	Glycerophosphocholines	PC
PC(18:1_22:5)	PC 18:1_22:5	833.5935	C48H84NO8P	Glycerophospholipids	Glycerophosphocholines	PC
PC(16:0_22:6)	PC 16:0_22:6	805.5622	C46H80NO8P	Glycerophospholipids	Glycerophosphocholines	PC
PC(16:1_22:6)	PC 16:1_22:6	803.5465	C46H78NO8P	Glycerophospholipids	Glycerophosphocholines	PC
PC(18:0_22:6)	PC 18:0_22:6	833.5935	C48H84NO8P	Glycerophospholipids	Glycerophosphocholines	PC
PC(18:1_22:6)	PC 18:1_22:6	831.5778	C48H82NO8P	Glycerophospholipids	Glycerophosphocholines	PC
PC(16:0_20:4-d1)	-			Glycerophospholipids	Glycerophosphocholines	PC
PC(16:1_20:4-d1)	-			Glycerophospholipids	Glycerophosphocholines	PC
PC(18:0_20:4-d1)	-			Glycerophospholipids	Glycerophosphocholines	PC
PC(18:1_20:4-d1)	-			Glycerophospholipids	Glycerophosphocholines	PC
PC(16:0_22:4-d1)	-			Glycerophospholipids	Glycerophosphocholines	PC
PC(16:0_22:4-d2)	-			Glycerophospholipids	Glycerophosphocholines	PC
PC(18:0_22:4-d1)	-			Glycerophospholipids	Glycerophosphocholines	PC
PC(18:0_22:4-d2)	-			Glycerophospholipids	Glycerophosphocholines	PC
PC(18:1_22:4-d1)	-			Glycerophospholipids	Glycerophosphocholines	PC
PC(18:1_22:4-d2)	-			Glycerophospholipids	Glycerophosphocholines	PC
PC(16:0_22:5-d1)	-			Glycerophospholipids	Glycerophosphocholines	PC
PC(16:0_22:5-d2)	-			Glycerophospholipids	Glycerophosphocholines	PC
PC(18:0_22:5-d1)	-			Glycerophospholipids	Glycerophosphocholines	PC
PC(18:0_22:5-d2)	-			Glycerophospholipids	Glycerophosphocholines	PC
PC(18:1_22:5-d1)	-			Glycerophospholipids	Glycerophosphocholines	PC
PC(18:1_22:5-d2)	-			Glycerophospholipids	Glycerophosphocholines	PC
PC(16:0_22:6-d1)	-			Glycerophospholipids	Glycerophosphocholines	PC
PC(16:0_22:6-d2)	-			Glycerophospholipids	Glycerophosphocholines	PC
PC(18:0_22:6-d1)	-			Glycerophospholipids	Glycerophosphocholines	PC
PC(18:0_22:6-d2)	-			Glycerophospholipids	Glycerophosphocholines	PC
PC(18:1_22:6-d1)	-			Glycerophospholipids	Glycerophosphocholines	PC
PC(18:1_22:6-d2)	-			Glycerophospholipids	Glycerophosphocholines	PC
METABOLITES_END
#END