#METABOLOMICS WORKBENCH Unok_20241119_004824 DATATRACK_ID:5381 STUDY_ID:ST003586 ANALYSIS_ID:AN005890 PROJECT_ID:PR002216 VERSION 1 CREATED_ON November 21, 2024, 8:35 pm #PROJECT PR:PROJECT_TITLE Metabolomics of the liver after administration of adenovirus associated with PR:PROJECT_TITLE metabolic liver zonation PR:PROJECT_SUMMARY Metabolic zonation in the liver is broadly defined as spatiotemporal variability PR:PROJECT_SUMMARY in the intra-hepatic distribution of nutrients. R-spondin3 (Rspo3) regulates PR:PROJECT_SUMMARY metabolic features around hepatic central veins. Herein, we demonstrate that PR:PROJECT_SUMMARY altered hepatic metabolic zonation due to Rspo3 contributes to maintaining PR:PROJECT_SUMMARY systemic glucose metabolism and body composition via inter-organ communication PR:PROJECT_SUMMARY mechanism. Hepatic Rspo3 induction in obesity improves obesity-associated PR:PROJECT_SUMMARY features by restoring insulin sensitivity, reversing adipose tissue enlargement PR:PROJECT_SUMMARY and reversing overstimulated adaptive thermogenesis. These remote effects partly PR:PROJECT_SUMMARY consist of neuronal communication via the afferent vagus from the liver. In PR:PROJECT_SUMMARY contrast, hepatic Rspo3 suppression exacerbates diabetes due to insulin PR:PROJECT_SUMMARY resistance and develops fatty liver. PR:INSTITUTE Teikyo University PR:LAST_NAME Uno PR:FIRST_NAME Kenji PR:ADDRESS 2-11-1 Kaga, Itabashi-ku, Tokyo PR:EMAIL unok@med.teikyo-u.ac.jp PR:PHONE 81339641211 #STUDY ST:STUDY_TITLE Metabolomics of the murine liver ST:STUDY_SUMMARY To examine whether a molecule associated with metabolic liver zonation has an ST:STUDY_SUMMARY effect on systemic metabolism, we first administer adenoviruses expressing Rspo3 ST:STUDY_SUMMARY or LacZ (control) into the murine liver. Then, by performing metabolome ST:STUDY_SUMMARY analyses, we compare the results of Rspo3-mice with those of LacZ-mice. ST:INSTITUTE Teikyo University ST:LAST_NAME Uno ST:FIRST_NAME Kenji ST:ADDRESS 2-11-1 Kaga, Itabashi-ku, Tokyo ST:EMAIL unok@med.teikyo-u.ac.jp ST:PHONE 81339641211 #SUBJECT SU:SUBJECT_TYPE Mammal SU:SUBJECT_SPECIES Mus musculus SU:TAXONOMY_ID 10090 #FACTORS #SUBJECT_SAMPLE_FACTORS: SUBJECT(optional)[tab]SAMPLE[tab]FACTORS(NAME:VALUE pairs separated by |)[tab]Raw file names and additional sample data SUBJECT_SAMPLE_FACTORS - ob-LacZ-1 Sample source:liver | Group:ob-LacZ SUBJECT_SAMPLE_FACTORS - ob-LacZ-2 Sample source:liver | Group:ob-LacZ SUBJECT_SAMPLE_FACTORS - ob-LacZ-3 Sample source:liver | Group:ob-LacZ SUBJECT_SAMPLE_FACTORS - ob-LacZ-4 Sample source:liver | Group:ob-LacZ SUBJECT_SAMPLE_FACTORS - ob-Rspo3-1 Sample source:liver | Group:ob-Rspo3 SUBJECT_SAMPLE_FACTORS - ob-Rspo3-2 Sample source:liver | Group:ob-Rspo3 SUBJECT_SAMPLE_FACTORS - ob-Rspo3-3 Sample source:liver | Group:ob-Rspo3 SUBJECT_SAMPLE_FACTORS - ob-Rspo3-4 Sample source:liver | Group:ob-Rspo3 #COLLECTION CO:COLLECTION_SUMMARY Liver tissues were rapidly removed on day 7 after adenovirus administration, CO:COLLECTION_SUMMARY frozen on liquid nitrogen and stored at –80°C until further processing. Then, CO:COLLECTION_SUMMARY liver samples were weighed at approximately 40mg and subjected to the experiment CO:COLLECTION_SUMMARY of metabolomics. CO:SAMPLE_TYPE Liver #TREATMENT TR:TREATMENT_SUMMARY Recombinant adenoviruses expressing LacZ or Rspo3 were administered into the TR:TREATMENT_SUMMARY liver of standard diet-fed ob/ob mice through tail vein injection. Liver tissues TR:TREATMENT_SUMMARY were taken on day 7 after adenovirus administration, subjected to the experiment TR:TREATMENT_SUMMARY of metabolomics. #SAMPLEPREP SP:SAMPLEPREP_SUMMARY Approximately 40 mg of frozen liver tissues were placed in a homogenization SP:SAMPLEPREP_SUMMARY tube, along with zirconia beads (5mmφ and 3mmφ). These tissues with internal SP:SAMPLEPREP_SUMMARY standards were homogenized with ice-cold acetonitrile:water (50:50) and SP:SAMPLEPREP_SUMMARY centrifuged at 2,300 × g for 5 min at 4 °C. The supernatant was filtered SP:SAMPLEPREP_SUMMARY through a 5-kDa cut-off filter (Ultrafree-MC-PLHCC, Human Metabolome SP:SAMPLEPREP_SUMMARY Technologies, Japan) at 9,100 × g for 180 min at 4 °C. The filtrate was SP:SAMPLEPREP_SUMMARY evaporated to dryness under vacuum and reconstituted in 50 µL of Milli-Q water SP:SAMPLEPREP_SUMMARY for metabolome analysis at HMT. #CHROMATOGRAPHY CH:CHROMATOGRAPHY_TYPE CE CH:INSTRUMENT_NAME Agilent 7100 CE CH:COLUMN_NAME Fused silica capillary i.d. 50 um x 80 cm CH:SOLVENT_A Anion Buffer Solution (p/n : I3302-1023) CH:SOLVENT_B N/A CH:FLOW_GRADIENT N/A CH:FLOW_RATE N/A CH:COLUMN_TEMPERATURE N/A #ANALYSIS AN:ANALYSIS_TYPE MS #MS MS:INSTRUMENT_NAME Agilent 6230 TOF MS:INSTRUMENT_TYPE TOF MS:MS_TYPE ESI MS:ION_MODE NEGATIVE MS:MS_COMMENTS The spectrometer was scanned from m/z 50 to 1,000. Peaks were extracted using MS:MS_COMMENTS automatic integration software MasterHands (ver. 2.19.0.2 Keio University, MS:MS_COMMENTS Japan). Signal peaks corresponding to isotopomers, adduct ions, and other MS:MS_COMMENTS product ions of known metabolites were excluded, and remaining peaks were MS:MS_COMMENTS annotated with putative metabolites from the HMT metabolite database based on MS:MS_COMMENTS their migration times (MTs) and m/z values. The tolerance range for the peak MS:MS_COMMENTS annotation was configured at ±0.5 min for MT and ±10 ppm for m/z. In addition, MS:MS_COMMENTS peak areas were normalized against those of the internal standards and then the MS:MS_COMMENTS resultant relative area values were further normalized by sample amount. #MS_METABOLITE_DATA MS_METABOLITE_DATA:UNITS arbitrary unit MS_METABOLITE_DATA_START Samples ob-LacZ-1 ob-LacZ-2 ob-LacZ-3 ob-LacZ-4 ob-Rspo3-1 ob-Rspo3-2 ob-Rspo3-3 ob-Rspo3-4 Factors Sample source:liver | Group:ob-LacZ Sample source:liver | Group:ob-LacZ Sample source:liver | Group:ob-LacZ Sample source:liver | Group:ob-LacZ Sample source:liver | Group:ob-Rspo3 Sample source:liver | Group:ob-Rspo3 Sample source:liver | Group:ob-Rspo3 Sample source:liver | Group:ob-Rspo3 Propionic acid 2.6E-04 N.D. N.D. N.D. 2.7E-04 3.1E-04 N.D. 3.0E-04 Pyruvic acid 3.8E-03 3.5E-03 3.5E-03 3.4E-03 3.0E-03 3.1E-03 4.3E-03 2.7E-03 Isobutyric acid" 1.5E-04 8.9E-05 1.2E-04 1.2E-04 2.9E-04 3.1E-04 2.1E-04 3.0E-04 Lactic acid 2.9E-01 2.4E-01 2.5E-01 2.1E-01 2.5E-01 2.5E-01 2.4E-01 2.2E-01 Valeric acid" N.D. N.D. N.D. N.D. 8.5E-05 8.0E-05 N.D. 9.7E-05 2-Hydroxyisobutyric acid N.D. N.D. N.D. 1.4E-04 1.9E-04 2.3E-04 2.2E-04 2.8E-04 2-Hydroxybutyric acid 5.9E-04 7.2E-04 8.7E-04 4.6E-04 8.3E-04 6.1E-04 4.4E-04 5.7E-04 3-Hydroxybutyric acid 3.0E-03 2.9E-03 2.6E-03 3.1E-03 4.5E-03 3.1E-03 4.1E-03 8.4E-03 Fumaric acid 2.0E-03 1.6E-03 1.2E-03 1.3E-03 3.9E-03 4.1E-03 3.7E-03 4.2E-03 Hexanoic acid 1.6E-04 1.4E-04 1.3E-04 1.4E-04 1.8E-04 2.1E-04 2.0E-04 1.8E-04 N-Acetylglycine N.D. N.D. N.D. 4.7E-05 9.6E-05 7.2E-05 9.2E-05 1.0E-04 Succinic acid 1.6E-02 1.9E-02 1.3E-02 1.8E-02 2.3E-02 1.7E-02 1.6E-02 1.6E-02 2-Hydroxyvaleric acid 2.3E-04 2.1E-04 1.8E-04 2.7E-04 6.7E-04 2.7E-04 3.2E-04 3.8E-04 Isethionic acid 2.0E-03 2.0E-03 2.7E-03 2.4E-03 8.3E-04 1.5E-03 9.4E-04 8.3E-04 5-Oxoproline 9.8E-04 8.4E-04 9.6E-04 1.3E-03 2.3E-03 2.6E-03 2.5E-03 3.8E-03 Itaconic acid N.D. N.D. N.D. N.D. 4.0E-04 3.4E-04 3.1E-04 4.4E-04 2-Oxohexanoic acid" N.D. N.D. N.D. N.D. N.D. N.D. N.D. 8.5E-05 Heptanoic acid 1.0E-04 4.0E-05 N.D. 5.3E-05 N.D. 6.9E-05 7.6E-05 5.4E-05 N-Acetylalanine 2.0E-04 1.3E-04 1.5E-04 2.0E-04 3.2E-04 4.0E-04 3.6E-04 5.8E-04 Methylsuccinic acid" 2.8E-04 N.D. 4.0E-04 N.D. N.D. 3.7E-04 N.D. N.D. 3-Ureidopropionic acid 2.9E-04 1.8E-04 2.1E-04 1.9E-04 7.6E-04 7.7E-04 6.1E-04 6.8E-04 Malic acid 1.2E-02 8.9E-03 4.8E-03 4.6E-03 2.3E-02 2.4E-02 2.0E-02 2.5E-02 m-Toluic acid" 1.7E-03 1.1E-03 1.0E-03 9.5E-04 1.1E-03 1.1E-03 1.5E-03 1.5E-03 Ethanolamine phosphate 2.2E-03 1.7E-03 1.8E-03 1.8E-03 5.3E-03 5.0E-03 4.8E-03 6.2E-03 N-Ethylmaleimide_+H2O N.D. N.D. N.D. N.D. N.D. N.D. N.D. N.D. Octanoic acid 1.4E-04 1.2E-04 1.1E-04 1.0E-04 1.2E-04 1.5E-04 1.6E-04 1.4E-04 4-Acetamidobutanoic acid 1.5E-04 2.2E-04 3.3E-04 1.8E-04 2.8E-04 3.8E-04 3.9E-04 2.1E-04 2-Hydroxyglutaric acid 1.0E-03 8.8E-04 8.4E-04 1.1E-03 2.3E-03 1.7E-03 2.0E-03 1.8E-03 Oxypurinol N.D. N.D. N.D. N.D. N.D. 6.9E-05 N.D. 1.3E-04 Pelargonic acid 3.7E-04 2.9E-04 2.9E-04 3.0E-04 2.9E-04 3.0E-04 3.2E-04 3.7E-04 Terephthalic acid 1.6E-04 1.3E-04 1.2E-04 1.2E-04 1.3E-04 1.2E-04 1.6E-04 1.4E-04 XA0012 1.2E-04 1.3E-04 1.6E-04 1.5E-04 2.1E-04 2.1E-04 1.7E-04 2.0E-04 Phosphoenolpyruvic acid N.D. N.D. N.D. N.D. N.D. N.D. N.D. N.D. Uric acid 8.9E-05 1.2E-04 7.0E-05 4.7E-05 1.9E-03 6.6E-04 6.0E-04 1.3E-03 Dihydroxyacetone phosphate 2.4E-03 7.4E-04 9.2E-04 1.1E-03 2.5E-03 3.2E-03 3.4E-03 3.6E-03 Glyceraldehyde 3-phosphate N.D. N.D. N.D. N.D. 7.4E-05 7.7E-05 6.6E-05 6.5E-05 Glycerol 3-phosphate 1.3E-02 1.6E-02 1.5E-02 1.5E-02 2.0E-02 1.5E-02 1.7E-02 1.7E-02 Decanoic acid 1.4E-04 1.0E-04 9.3E-05 1.6E-04 1.8E-04 1.3E-04 1.6E-04 1.2E-04 N-Acetylleucine" 1.3E-04 1.3E-04 1.6E-04 1.1E-04 1.1E-04 1.2E-04 1.5E-04 1.1E-04 cis-Aconitic acid 1.2E-04 9.2E-05 N.D. N.D. 3.1E-04 2.3E-04 2.5E-04 3.3E-04 N-Acetylasparagine N.D. 7.9E-05 8.9E-05 N.D. 1.5E-04 9.8E-05 9.9E-05 N.D. Formiminoglutamic acid N.D. N.D. N.D. N.D. 1.1E-04 6.4E-05 9.6E-05 5.4E-05 N-Acetylaspartic acid N.D. N.D. N.D. N.D. 2.3E-04 1.4E-04 1.1E-04 1.4E-04 Ascorbic acid 1.6E-02 1.3E-02 1.4E-02 1.3E-02 3.3E-02 3.9E-02 3.5E-02 3.6E-02 Hippuric acid 5.5E-05 7.3E-05 N.D. 6.9E-05 1.1E-04 5.0E-05 N.D. N.D. Hydroxyphenyllactic acid" 2.3E-04 1.9E-04 2.8E-04 2.2E-04 1.1E-04 1.1E-04 1.3E-04 1.5E-04 3-Phosphoglyceric acid 1.7E-04 1.8E-04 1.4E-04 9.5E-05 2.1E-04 1.7E-04 2.1E-04 2.1E-04 XA0017 1.4E-04 1.0E-04 1.3E-04 1.2E-04 1.1E-04 7.8E-05 1.5E-04 1.0E-04 N-Acetylglutamine 6.3E-04 9.6E-04 7.4E-04 6.9E-04 1.5E-03 1.3E-03 1.4E-03 1.1E-03 Azelaic acid 7.5E-05 6.8E-05 N.D. N.D. 7.1E-05 9.5E-05 N.D. 7.3E-05 N-Acetylglutamic acid 7.6E-04 9.0E-04 1.1E-03 1.2E-03 2.4E-03 1.7E-03 2.5E-03 1.7E-03 N-Acetylmethionine 1.8E-04 1.2E-04 1.1E-04 1.3E-04 3.6E-04 2.8E-04 2.4E-04 4.3E-04 XA0019 1.5E-03 2.4E-03 1.6E-03 2.2E-03 3.2E-03 3.1E-03 3.6E-03 3.1E-03 Citric acid 2.6E-03 1.8E-03 1.7E-03 1.6E-03 6.5E-03 4.5E-03 5.4E-03 7.5E-03 Phenaceturic acid N.D. N.D. N.D. N.D. N.D. N.D. N.D. N.D. Glucuronic acid-1" 1.8E-04 2.9E-04 2.3E-04 2.2E-04 6.1E-04 5.4E-04 6.5E-04 6.0E-04 Glucuronic acid-2" 1.6E-03 1.6E-03 1.4E-03 1.6E-03 1.4E-03 1.8E-03 1.7E-03 2.2E-03 Gluconic acid 7.8E-03 8.7E-03 6.6E-03 6.2E-03 1.5E-02 1.2E-02 1.5E-02 1.5E-02 Lauric acid 8.9E-04 5.7E-04 5.8E-04 7.0E-04 6.5E-04 5.9E-04 6.4E-04 6.3E-04 "6,8-Thioctic acid" N.D. N.D. N.D. N.D. 6.5E-05 7.0E-05 5.7E-05 6.3E-05 Glucaric acid 8.7E-05 8.0E-05 7.3E-05 6.0E-05 7.3E-05 N.D. 1.1E-04 N.D. Mucic acid 9.8E-05 1.6E-04 1.9E-04 1.9E-04 1.9E-04 2.3E-04 2.9E-04 2.3E-04 3-Indoxylsulfuric acid N.D. 6.0E-05 5.1E-05 4.8E-05 9.8E-05 N.D. 1.0E-04 8.3E-05 Pantothenic acid 2.6E-04 2.1E-04 2.6E-04 2.4E-04 2.4E-04 3.5E-04 2.9E-04 4.3E-04 Myristoleic acid 2.3E-04 2.0E-04 2.2E-04 2.6E-04 2.1E-04 2.4E-04 2.0E-04 2.3E-04 Myristic acid 5.4E-04 4.6E-04 5.0E-04 5.3E-04 4.9E-04 6.3E-04 4.0E-04 4.8E-04 Ribulose 5-phosphate 8.3E-03 8.1E-03 8.7E-03 8.0E-03 8.1E-03 8.2E-03 9.3E-03 8.8E-03 Ribose 5-phosphate 9.6E-04 8.4E-04 7.4E-04 1.0E-03 1.4E-03 1.1E-03 1.6E-03 1.2E-03 XA0033 4.9E-03 2.8E-03 5.3E-03 2.5E-03 8.4E-03 7.6E-03 7.6E-03 8.9E-03 -Glu-Taurine 1.1E-04 6.8E-05 9.5E-05 1.1E-04 1.7E-04 2.6E-04 1.3E-04 2.5E-04 Ascorbate 2-sulfate" N.D. N.D. N.D. N.D. 8.3E-05 9.1E-05 1.2E-04 N.D. XA0035 2.9E-04 3.0E-04 2.8E-04 2.3E-04 2.9E-04 3.1E-04 3.9E-04 2.8E-04 Ascorbate 2-phosphate 1.2E-04 1.7E-04 9.7E-05 1.1E-04 1.1E-04 N.D. 1.1E-04 1.1E-04 myo-Inositol 3-phosphate" 5.0E-04 4.8E-04 5.2E-04 4.9E-04 4.2E-04 3.5E-04 4.5E-04 5.1E-04 Glucose 6-phosphate 1.7E-02 1.5E-02 1.3E-02 1.6E-02 2.5E-02 2.6E-02 2.7E-02 3.1E-02 myo-Inositol 2-phosphate 3.2E-04 3.5E-04 3.2E-04 2.9E-04 3.4E-04 4.5E-04 3.9E-04 5.0E-04 Fructose 6-phosphate 3.8E-03 3.3E-03 2.8E-03 3.7E-03 6.5E-03 6.8E-03 6.8E-03 7.9E-03 Glucose 1-phosphate 3.4E-03 3.0E-03 2.7E-03 3.0E-03 3.2E-03 3.7E-03 3.5E-03 4.1E-03 "2,3-Diphosphoglyceric acid" 1.3E-03 1.6E-03 1.6E-03 1.2E-03 3.8E-03 2.0E-03 2.9E-03 3.0E-03 6-Phosphogluconic acid 7.8E-04 7.6E-04 6.3E-04 6.3E-04 9.0E-04 7.3E-04 7.2E-04 9.1E-04 Xanthosine 3.1E-04 2.8E-04 2.9E-04 1.8E-04 3.4E-04 3.7E-04 3.0E-04 4.3E-04 Orotidine N.D. N.D. N.D. N.D. 4.1E-05 4.2E-05 6.1E-05 N.D. Sedoheptulose 7-phosphate 7.1E-03 6.5E-03 5.6E-03 6.3E-03 8.7E-03 8.1E-03 7.8E-03 9.6E-03 N-Acetylglucosamine 6-phosphate 1.5E-04 1.4E-04 1.7E-04 1.5E-04 2.6E-04 3.0E-04 2.9E-04 3.6E-04 N-Acetylglucosamine 1-phosphate 1.6E-04 1.5E-04 1.7E-04 1.1E-04 1.4E-04 1.7E-04 2.3E-04 2.2E-04 N-Acetylneuraminic acid 1.2E-04 1.2E-04 1.2E-04 1.3E-04 2.3E-04 1.9E-04 2.0E-04 2.3E-04 "Ribulose 1,5-diphosphate" 9.9E-05 7.9E-05 N.D. N.D. 1.2E-04 N.D. 1.0E-04 N.D. CMP 1.2E-03 1.4E-03 1.4E-03 1.5E-03 1.3E-03 1.3E-03 1.7E-03 1.4E-03 UMP 9.9E-03 9.0E-03 9.6E-03 9.1E-03 8.7E-03 9.0E-03 1.1E-02 1.1E-02 N-Glycolylneuraminic acid 6.9E-04 6.1E-04 4.2E-04 5.2E-04 1.1E-03 1.0E-03 9.5E-04 1.0E-03 5-Aminoimidazole-4-carboxamide ribotide 7.4E-05 6.5E-05 8.3E-05 4.5E-05 N.D. 5.6E-05 5.9E-05 N.D. Ascorbate 2-glucoside 1.8E-04 1.6E-04 1.7E-04 1.9E-04 2.4E-04 3.9E-04 3.2E-04 3.5E-04 "Fructose 1,6-diphosphate" 8.8E-04 2.8E-04 3.5E-04 4.0E-04 2.2E-03 3.2E-03 2.7E-03 3.9E-03 AMP 6.3E-02 6.9E-02 5.9E-02 6.9E-02 6.6E-02 6.7E-02 7.1E-02 5.3E-02 IMP 7.3E-03 6.8E-03 8.7E-03 5.7E-03 5.3E-03 6.2E-03 8.2E-03 1.2E-02 GMP 9.0E-03 9.0E-03 8.4E-03 9.4E-03 9.8E-03 1.0E-02 1.1E-02 9.9E-03 XA0055 4.2E-04 N.D. 1.5E-04 1.9E-04 9.3E-04 1.4E-03 1.1E-03 1.9E-03 NADPH_divalent 1.3E-04 1.2E-04 1.2E-04 1.7E-04 N.D. N.D. 1.0E-04 N.D. CoA_divalent 4.7E-03 4.3E-03 4.8E-03 4.7E-03 3.5E-03 3.2E-03 4.6E-03 2.9E-03 FAD_divalent 5.8E-04 6.3E-04 6.3E-04 7.2E-04 6.0E-04 5.6E-04 6.2E-04 6.0E-04 CDP N.D. N.D. 6.5E-05 5.8E-05 4.3E-05 N.D. N.D. N.D. UDP 2.6E-04 2.4E-04 2.4E-04 2.6E-04 2.2E-04 1.6E-04 2.0E-04 1.5E-04 Acetyl CoA_divalent 1.0E-04 1.3E-04 1.2E-04 6.8E-05 7.3E-05 N.D. N.D. N.D. Cholic acid 1.2E-04 2.3E-04 1.6E-04 2.8E-04 1.6E-03 2.5E-04 1.4E-03 1.7E-03 Isobutyryl CoA_divalent 8.9E-05 6.3E-05 7.0E-05 8.1E-05 9.3E-05 6.1E-05 6.1E-05 N.D. Thiamine diphosphate 1.3E-04 1.5E-04 8.7E-05 7.9E-05 4.8E-05 N.D. N.D. N.D. Adenosine 5'-phosphosulfate 6.3E-05 5.5E-05 6.3E-05 5.7E-05 N.D. N.D. N.D. N.D. ADP 1.1E-02 1.2E-02 1.2E-02 1.1E-02 7.2E-03 6.8E-03 9.0E-03 6.0E-03 GDP 4.9E-04 5.5E-04 4.7E-04 4.9E-04 6.2E-04 5.7E-04 6.0E-04 5.8E-04 XA0065 9.5E-04 9.4E-04 1.2E-03 8.6E-04 9.0E-04 8.5E-04 9.5E-04 1.2E-03 Adenylosuccinic acid 1.5E-03 1.5E-03 1.4E-03 1.5E-03 2.9E-03 3.4E-03 3.3E-03 3.0E-03 Glycocholic acid N.D. N.D. 3.4E-05 4.9E-05 6.1E-05 N.D. N.D. N.D. CDP-choline 4.8E-04 5.2E-04 5.2E-04 3.5E-04 4.3E-04 5.6E-04 4.2E-04 5.9E-04 ATP 1.3E-03 1.5E-03 1.7E-03 1.6E-03 7.7E-04 5.2E-04 1.0E-03 5.7E-04 Taurocholic acid 4.1E-02 5.1E-02 4.2E-02 5.5E-02 6.1E-02 3.2E-02 2.8E-02 4.0E-02 GTP 6.5E-05 6.3E-05 N.D. 5.7E-05 N.D. N.D. N.D. N.D. ADP-ribose 4.5E-03 6.1E-03 4.1E-03 3.4E-03 2.0E-03 8.7E-04 1.8E-03 1.8E-04 dTDP-glucose 1.1E-04 9.6E-05 1.2E-04 1.0E-04 1.2E-04 1.4E-04 1.6E-04 1.6E-04 UDP-glucose" 1.3E-02 1.3E-02 1.1E-02 1.2E-02 2.1E-02 2.3E-02 1.9E-02 1.9E-02 UDP-glucuronic acid 1.2E-02 1.5E-02 1.0E-02 9.9E-03 1.6E-02 1.1E-02 1.5E-02 1.4E-02 ADP-glucose" 2.1E-04 1.9E-04 1.8E-04 2.1E-04 2.9E-04 3.0E-04 3.1E-04 3.3E-04 GDP-glucose" 4.9E-04 5.3E-04 4.3E-04 4.7E-04 4.6E-04 4.8E-04 4.1E-04 3.7E-04 UDP-N-acetylglucosamine" 6.1E-03 6.0E-03 6.8E-03 7.3E-03 9.7E-03 9.6E-03 1.0E-02 1.2E-02 CMP-N-acetylneuraminate 5.1E-05 5.5E-05 N.D. 4.7E-05 1.4E-04 1.1E-04 1.4E-04 1.5E-04 NAD+ 1.5E-03 3.4E-03 2.0E-03 2.0E-03 9.4E-04 5.9E-04 8.6E-04 4.6E-04 3'-Dephospho CoA 1.1E-03 8.0E-04 1.0E-03 1.0E-03 1.2E-03 1.3E-03 1.4E-03 1.4E-03 NADP+ 5.7E-04 9.1E-04 7.7E-04 8.1E-04 7.5E-04 6.7E-04 1.1E-03 6.1E-04 MS_METABOLITE_DATA_END #METABOLITES METABOLITES_START metabolite_name Propionic acid Pyruvic acid Isobutyric acid" Lactic acid Valeric acid" 2-Hydroxyisobutyric acid 2-Hydroxybutyric acid 3-Hydroxybutyric acid Fumaric acid Hexanoic acid N-Acetylglycine Succinic acid 2-Hydroxyvaleric acid Isethionic acid 5-Oxoproline Itaconic acid 2-Oxohexanoic acid" Heptanoic acid N-Acetylalanine Methylsuccinic acid" 3-Ureidopropionic acid Malic acid m-Toluic acid" Ethanolamine phosphate N-Ethylmaleimide_+H2O Octanoic acid 4-Acetamidobutanoic acid 2-Hydroxyglutaric acid Oxypurinol Pelargonic acid Terephthalic acid XA0012 Phosphoenolpyruvic acid Uric acid Dihydroxyacetone phosphate Glyceraldehyde 3-phosphate Glycerol 3-phosphate Decanoic acid N-Acetylleucine" cis-Aconitic acid N-Acetylasparagine Formiminoglutamic acid N-Acetylaspartic acid Ascorbic acid Hippuric acid Hydroxyphenyllactic acid" 3-Phosphoglyceric acid XA0017 N-Acetylglutamine Azelaic acid N-Acetylglutamic acid N-Acetylmethionine XA0019 Citric acid Phenaceturic acid Glucuronic acid-1" Glucuronic acid-2" Gluconic acid Lauric acid "6,8-Thioctic acid" Glucaric acid Mucic acid 3-Indoxylsulfuric acid Pantothenic acid Myristoleic acid Myristic acid Ribulose 5-phosphate Ribose 5-phosphate XA0033 -Glu-Taurine Ascorbate 2-sulfate" XA0035 Ascorbate 2-phosphate myo-Inositol 3-phosphate" Glucose 6-phosphate myo-Inositol 2-phosphate Fructose 6-phosphate Glucose 1-phosphate "2,3-Diphosphoglyceric acid" 6-Phosphogluconic acid Xanthosine Orotidine Sedoheptulose 7-phosphate N-Acetylglucosamine 6-phosphate N-Acetylglucosamine 1-phosphate N-Acetylneuraminic acid "Ribulose 1,5-diphosphate" CMP UMP N-Glycolylneuraminic acid 5-Aminoimidazole-4-carboxamide ribotide Ascorbate 2-glucoside "Fructose 1,6-diphosphate" AMP IMP GMP XA0055 NADPH_divalent CoA_divalent FAD_divalent CDP UDP Acetyl CoA_divalent Cholic acid Isobutyryl CoA_divalent Thiamine diphosphate Adenosine 5'-phosphosulfate ADP GDP XA0065 Adenylosuccinic acid Glycocholic acid CDP-choline ATP Taurocholic acid GTP ADP-ribose dTDP-glucose UDP-glucose" UDP-glucuronic acid ADP-glucose" GDP-glucose" UDP-N-acetylglucosamine" CMP-N-acetylneuraminate NAD+ 3'-Dephospho CoA NADP+ METABOLITES_END #END