#METABOLOMICS WORKBENCH juliehaines_20241125_131224 DATATRACK_ID:5396 STUDY_ID:ST003604 ANALYSIS_ID:AN005922 PROJECT_ID:PR002231 VERSION 1 CREATED_ON December 5, 2024, 7:19 pm #PROJECT PR:PROJECT_TITLE Macrophages recycle phagocytosed bacteria to fuel immunometabolic responses PR:PROJECT_SUMMARY Macrophages specialize in phagocytosis, a cellular process that eliminates PR:PROJECT_SUMMARY extracellular matter, including microbes, through internalization and PR:PROJECT_SUMMARY degradation. Despite the critical role of phagocytosis during bacterial PR:PROJECT_SUMMARY infection, the fate of phagocytosed microbial cargo and its impact on host cell PR:PROJECT_SUMMARY is poorly understood. Here, we reveal that ingested bacteria constitute an PR:PROJECT_SUMMARY alternative nutrient source that skews immunometabolic host responses. Tracing PR:PROJECT_SUMMARY stable isotope-labelled bacteria, we found that phagolysosomal degradation of PR:PROJECT_SUMMARY bacteria provides carbon atoms and amino acids that are recycled into various PR:PROJECT_SUMMARY metabolic pathways, including glutathione and itaconate biosynthesis, and PR:PROJECT_SUMMARY satisfy macrophage bioenergetic needs. Metabolic recycling of PR:PROJECT_SUMMARY microbially-derived nutrients is regulated by the nutrient sensing mTORC1 and PR:PROJECT_SUMMARY intricately tied to microbial viability. Dead bacteria, as opposed to live ones, PR:PROJECT_SUMMARY are enriched in cyclic- adenosine monophosphate (AMP), sustain the cellular AMP PR:PROJECT_SUMMARY pool and subsequently activate AMP protein kinase (AMPK) to inhibit mTORC1. PR:PROJECT_SUMMARY Consequently, killed bacteria strongly fuel metabolic recycling and support PR:PROJECT_SUMMARY macrophage survival, but elicit decreased reactive oxygen species (ROS) PR:PROJECT_SUMMARY production and a reduced IL-1β secretion compared to viable bacteria. These PR:PROJECT_SUMMARY results reveal a novel insight into the fate of engulfed microbes and highlights PR:PROJECT_SUMMARY a microbial viability-associated metabolite that triggers host metabolic and PR:PROJECT_SUMMARY immune responses. Our findings hold promise for shaping immunometabolic PR:PROJECT_SUMMARY intervention in various immune-related pathologies. PR:INSTITUTE University of Colorado Anschutz Medical Campus PR:LABORATORY Lab of Angelo D'Alessandro in collaboration with lab of Johan Garaude (INSERM, PR:LABORATORY Fr) PR:LAST_NAME Haines PR:FIRST_NAME Julie PR:ADDRESS 12801 E 17th Ave, Room 1303, Aurora, Colorado, 80045, USA PR:EMAIL julie.haines@cuanschutz.edu PR:PHONE 3037243339 #STUDY ST:STUDY_TITLE Investigation of global metabolites and E coli-derived metabolites following ST:STUDY_TITLE engulfment of dead bacteria by wild type or RagAGTP-expressing bone marrow ST:STUDY_TITLE derived macrophages. ST:STUDY_SUMMARY Global metabolomics and 13C-tracing analysis of wild type or RagAGTP-expressing ST:STUDY_SUMMARY bone marrow derived macrophages exposed to heat-killed uniformly 13C-labelled E. ST:STUDY_SUMMARY coli in a trans-well system for 6h or 18h. ST:INSTITUTE University of Colorado Anschutz Medical Campus ST:LAST_NAME Haines ST:FIRST_NAME Julie ST:ADDRESS 12801 E 17th Ave, Room 1303, Aurora, Colorado, 80045, USA ST:EMAIL julie.haines@cuanschutz.edu ST:PHONE 3037243339 #SUBJECT SU:SUBJECT_TYPE Cultured cells SU:SUBJECT_SPECIES Mus musculus SU:GENDER Not applicable #SUBJECT_SAMPLE_FACTORS: SUBJECT(optional)[tab]SAMPLE[tab]FACTORS(NAME:VALUE pairs separated by |)[tab]Raw file names and additional sample data SUBJECT_SAMPLE_FACTORS - CT WT 1 time (h):0 | Dead E coli treatment:no | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:WT RAW_FILE_NAME(raw file name)=F2-64-01- SUBJECT_SAMPLE_FACTORS - CT WT 2 time (h):0 | Dead E coli treatment:no | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:WT RAW_FILE_NAME(raw file name)=F2-64-02- SUBJECT_SAMPLE_FACTORS - CT WT 3 time (h):0 | Dead E coli treatment:no | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:WT RAW_FILE_NAME(raw file name)=F2-64-03- SUBJECT_SAMPLE_FACTORS - CT WT 4 time (h):0 | Dead E coli treatment:no | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:WT RAW_FILE_NAME(raw file name)=F2-64-04- SUBJECT_SAMPLE_FACTORS - WT 1 6h time (h):6 | Dead E coli treatment:yes | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:WT RAW_FILE_NAME(raw file name)=F2-64-05- SUBJECT_SAMPLE_FACTORS - WT 2 6h time (h):6 | Dead E coli treatment:yes | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:WT RAW_FILE_NAME(raw file name)=F2-64-06- SUBJECT_SAMPLE_FACTORS - WT 3 6h time (h):6 | Dead E coli treatment:yes | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:WT RAW_FILE_NAME(raw file name)=F2-64-07- SUBJECT_SAMPLE_FACTORS - WT 4 6h time (h):6 | Dead E coli treatment:yes | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:WT RAW_FILE_NAME(raw file name)=F2-64-08- SUBJECT_SAMPLE_FACTORS - WT 1 18h time (h):18 | Dead E coli treatment:yes | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:WT RAW_FILE_NAME(raw file name)=F2-64-09- SUBJECT_SAMPLE_FACTORS - WT 2 18h time (h):18 | Dead E coli treatment:yes | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:WT RAW_FILE_NAME(raw file name)=F2-64-10- SUBJECT_SAMPLE_FACTORS - WT3 18h time (h):18 | Dead E coli treatment:yes | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:WT RAW_FILE_NAME(raw file name)=F2-64-11- SUBJECT_SAMPLE_FACTORS - WT 4 18h time (h):18 | Dead E coli treatment:yes | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:WT RAW_FILE_NAME(raw file name)=F2-64-12- SUBJECT_SAMPLE_FACTORS - CT RagA GTP 1 time (h):0 | Dead E coli treatment:no | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:RagA GTP mutant RAW_FILE_NAME(raw file name)=F2-64-13- SUBJECT_SAMPLE_FACTORS - CT RagA GTP 2 time (h):0 | Dead E coli treatment:no | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:RagA GTP mutant RAW_FILE_NAME(raw file name)=F2-64-14- SUBJECT_SAMPLE_FACTORS - CT RagA GTP 3 time (h):0 | Dead E coli treatment:no | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:RagA GTP mutant RAW_FILE_NAME(raw file name)=F2-64-15- SUBJECT_SAMPLE_FACTORS - CT RagA GTP 4 time (h):0 | Dead E coli treatment:no | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:RagA GTP mutant RAW_FILE_NAME(raw file name)=F2-64-16- SUBJECT_SAMPLE_FACTORS - RagA GTP 1 6h time (h):6 | Dead E coli treatment:yes | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:RagA GTP mutant RAW_FILE_NAME(raw file name)=F2-64-17- SUBJECT_SAMPLE_FACTORS - RagA GTP 2 6h time (h):6 | Dead E coli treatment:yes | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:RagA GTP mutant RAW_FILE_NAME(raw file name)=F2-64-18- SUBJECT_SAMPLE_FACTORS - RagA GTP 3 6h time (h):6 | Dead E coli treatment:yes | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:RagA GTP mutant RAW_FILE_NAME(raw file name)=F2-64-19- SUBJECT_SAMPLE_FACTORS - RagA GTP 4 6h time (h):6 | Dead E coli treatment:yes | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:RagA GTP mutant RAW_FILE_NAME(raw file name)=F2-64-20- SUBJECT_SAMPLE_FACTORS - RagA GTP 1 18h time (h):18 | Dead E coli treatment:yes | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:RagA GTP mutant RAW_FILE_NAME(raw file name)=F2-64-21- SUBJECT_SAMPLE_FACTORS - RagA GTP 2 18h time (h):18 | Dead E coli treatment:yes | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:RagA GTP mutant RAW_FILE_NAME(raw file name)=F2-64-22- SUBJECT_SAMPLE_FACTORS - RagA GTP 3 18h time (h):18 | Dead E coli treatment:yes | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:RagA GTP mutant RAW_FILE_NAME(raw file name)=F2-64-23- SUBJECT_SAMPLE_FACTORS - RagA GTP 4 18h time (h):18 | Dead E coli treatment:yes | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:RagA GTP mutant RAW_FILE_NAME(raw file name)=F2-64-24- SUBJECT_SAMPLE_FACTORS - CT WT 1 + Conca time (h):0 | Dead E coli treatment:no | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:WT RAW_FILE_NAME(raw file name)=F2-64-25- SUBJECT_SAMPLE_FACTORS - CT WT 2 + Conca time (h):0 | Dead E coli treatment:no | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:WT RAW_FILE_NAME(raw file name)=F2-64-26- SUBJECT_SAMPLE_FACTORS - CT WT 3 + Conca time (h):0 | Dead E coli treatment:no | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:WT RAW_FILE_NAME(raw file name)=F2-64-27- SUBJECT_SAMPLE_FACTORS - CT WT 4 + Conca time (h):0 | Dead E coli treatment:no | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:WT RAW_FILE_NAME(raw file name)=F2-64-28- SUBJECT_SAMPLE_FACTORS - WT 1 6h + Conca time (h):6 | Dead E coli treatment:yes | inhibitor:150 nM concanamycin A | Sample source:murine bone marrow derived macrophages | genotype:WT RAW_FILE_NAME(raw file name)=F2-64-29- SUBJECT_SAMPLE_FACTORS - WT 2 6h + Conca time (h):6 | Dead E coli treatment:yes | inhibitor:150 nM concanamycin A | Sample source:murine bone marrow derived macrophages | genotype:WT RAW_FILE_NAME(raw file name)=F2-64-30- SUBJECT_SAMPLE_FACTORS - WT 3 6h + Conca time (h):6 | Dead E coli treatment:yes | inhibitor:150 nM concanamycin A | Sample source:murine bone marrow derived macrophages | genotype:WT RAW_FILE_NAME(raw file name)=F2-64-31- SUBJECT_SAMPLE_FACTORS - WT 4 6h + Conca time (h):6 | Dead E coli treatment:yes | inhibitor:150 nM concanamycin A | Sample source:murine bone marrow derived macrophages | genotype:WT RAW_FILE_NAME(raw file name)=F2-64-32- SUBJECT_SAMPLE_FACTORS - WT 1 18h + Conca time (h):18 | Dead E coli treatment:yes | inhibitor:150 nM concanamycin A | Sample source:murine bone marrow derived macrophages | genotype:WT RAW_FILE_NAME(raw file name)=F2-64-33- SUBJECT_SAMPLE_FACTORS - WT 2 18h + Conca time (h):18 | Dead E coli treatment:yes | inhibitor:150 nM concanamycin A | Sample source:murine bone marrow derived macrophages | genotype:WT RAW_FILE_NAME(raw file name)=F2-64-34- SUBJECT_SAMPLE_FACTORS - WT3 18h + Conca time (h):18 | Dead E coli treatment:yes | inhibitor:150 nM concanamycin A | Sample source:murine bone marrow derived macrophages | genotype:WT RAW_FILE_NAME(raw file name)=F2-64-35- SUBJECT_SAMPLE_FACTORS - WT 4 18h + Conca time (h):18 | Dead E coli treatment:yes | inhibitor:150 nM concanamycin A | Sample source:murine bone marrow derived macrophages | genotype:WT RAW_FILE_NAME(raw file name)=F2-64-36- SUBJECT_SAMPLE_FACTORS - WT 1 6h + Bafilo time (h):6 | Dead E coli treatment:yes | inhibitor:50 nM bafilomycin A1 | Sample source:murine bone marrow derived macrophages | genotype:WT RAW_FILE_NAME(raw file name)=F2-64-37- SUBJECT_SAMPLE_FACTORS - WT 2 6h + Bafilo time (h):6 | Dead E coli treatment:yes | inhibitor:50 nM bafilomycin A1 | Sample source:murine bone marrow derived macrophages | genotype:WT RAW_FILE_NAME(raw file name)=F2-64-38- SUBJECT_SAMPLE_FACTORS - WT 3 6h + Bafilo time (h):6 | Dead E coli treatment:yes | inhibitor:50 nM bafilomycin A1 | Sample source:murine bone marrow derived macrophages | genotype:WT RAW_FILE_NAME(raw file name)=F2-64-39- SUBJECT_SAMPLE_FACTORS - WT 4 6h + Bafilo time (h):6 | Dead E coli treatment:yes | inhibitor:50 nM bafilomycin A1 | Sample source:murine bone marrow derived macrophages | genotype:WT RAW_FILE_NAME(raw file name)=F2-64-40- SUBJECT_SAMPLE_FACTORS - WT 1 18h + Bafilo time (h):18 | Dead E coli treatment:yes | inhibitor:50 nM bafilomycin A1 | Sample source:murine bone marrow derived macrophages | genotype:WT RAW_FILE_NAME(raw file name)=F2-64-41- SUBJECT_SAMPLE_FACTORS - WT 2 18h + Bafilo time (h):18 | Dead E coli treatment:yes | inhibitor:50 nM bafilomycin A1 | Sample source:murine bone marrow derived macrophages | genotype:WT RAW_FILE_NAME(raw file name)=F2-64-42- SUBJECT_SAMPLE_FACTORS - WT3 18h + Bafilo time (h):18 | Dead E coli treatment:yes | inhibitor:50 nM bafilomycin A1 | Sample source:murine bone marrow derived macrophages | genotype:WT RAW_FILE_NAME(raw file name)=F2-64-43- SUBJECT_SAMPLE_FACTORS - WT 4 18h + Bafilo time (h):18 | Dead E coli treatment:yes | inhibitor:50 nM bafilomycin A1 | Sample source:murine bone marrow derived macrophages | genotype:WT RAW_FILE_NAME(raw file name)=F2-64-44- #COLLECTION CO:COLLECTION_SUMMARY BMDMs were seeded at 2.5E6 cell/well in a non-tissue treated 6-well plate (2 CO:COLLECTION_SUMMARY wells per condition) the day before the experiment. To harvest, BMDMs were CO:COLLECTION_SUMMARY washed with cold PBS and harvested with 5 mM EDTA in PBS, frozen as dry cell CO:COLLECTION_SUMMARY pellet, and stored at -80˚C until processing. CO:SAMPLE_TYPE Macrophages #TREATMENT TR:TREATMENT_SUMMARY Preparation of macrophages: Murine bone marrow-derived macrophages (BMDMs) were TR:TREATMENT_SUMMARY generated as described previously, in RPMI 1640 supplemented with M-CSF (30% TR:TREATMENT_SUMMARY mycoplasma-free L929 cell supernatant, NCBI Biosample accession # SAMN00155972) TR:TREATMENT_SUMMARY and 10% FBS, plus 100 µg/ml penicillin, 100 µg/ml streptomycin, 10 mM HEPES, 1 TR:TREATMENT_SUMMARY nM sodium pyruvate and 50 µM 2-mercaptoethanol (all from Gibco). BMDMs were TR:TREATMENT_SUMMARY used on day 5 to 7 after seeding. Period of differentiation of the cells, TR:TREATMENT_SUMMARY concentration of cells when replating and time-lapse between replating and TR:TREATMENT_SUMMARY stimulation with bacteria were important parameters to maintain metabolic TR:TREATMENT_SUMMARY backgrounds homogenous between experiments. Preparation of viable and killed TR:TREATMENT_SUMMARY U-[13C]Bacteria: ThyA- E. coli were grown overnight with shaking in LB TR:TREATMENT_SUMMARY supplemented with thymidine (500 µg/ml) and trimethoprim (50 µg/ml), diluted TR:TREATMENT_SUMMARY 1/40, and grown until log-phase [optical density at 600 nm (OD600) of 0.8-1.2]. TR:TREATMENT_SUMMARY Bacteria were washed with phosphate buffer saline (PBS) to remove LB salts TR:TREATMENT_SUMMARY before addition to cells. For labeling of bacteria, 10 µl of an overnight TR:TREATMENT_SUMMARY cultured of thyA- E. coli was added to 20 ml of a filtered M9 minimal medium TR:TREATMENT_SUMMARY salts (Life Technologies) supplemented with 1 mM thiamine, 1 mM MgSO4, 0.1 M TR:TREATMENT_SUMMARY CaCl2, 500 µg/ml thymidine, 50 µg/ml trimethoprim, and 0.5% U-[13C6] glucose TR:TREATMENT_SUMMARY (Campro Scientific). Bacteria were grown for 72h, washed with PBS and subjected TR:TREATMENT_SUMMARY to heat-killing by re-suspension in PBS and subsequently incubation at 60˚C for TR:TREATMENT_SUMMARY 60-90 min. For antibiotic killing, bacteria were incubated for 6h to 12h with TR:TREATMENT_SUMMARY Streptomycin or Gentamycin (50 µg/ml). Bacteria were kept at 4˚C until use. TR:TREATMENT_SUMMARY Efficient killing was confirmed by overnight plating on LB-agar plates. TR:TREATMENT_SUMMARY Treatment of macrophages: 2E6 BMDMs were plated 12-16h prior stimulation in TR:TREATMENT_SUMMARY non-tissue cultured treated 6-well plate (BD Falcons). Cells were then TR:TREATMENT_SUMMARY stimulated with killed labelled-E. coli at MOI 50, centrifuged at 2000 rpm for TR:TREATMENT_SUMMARY 5min. BMDMs were incubated for 5 min and washed with PBS to remove non-ingested TR:TREATMENT_SUMMARY bacteria and further incubated for 6h or 18h. To inhibit lysosomal degradation, TR:TREATMENT_SUMMARY cells were treated with 50 nM Bafilomycin A1. To inhibit v-ATPase, cells were TR:TREATMENT_SUMMARY treated with 150 nM concanamycin A. All the above inhibitors were added to the TR:TREATMENT_SUMMARY cultured medium at least 2h before stimulation of the BMDMs. #SAMPLEPREP SP:SAMPLEPREP_SUMMARY Metabolites from frozen pellets were extracted at 4e6 cells per mL using ice SP:SAMPLEPREP_SUMMARY cold 5:3:2 methanol:acetonitrile:water (v/v/v) with vigorous vortexing at 4 SP:SAMPLEPREP_SUMMARY degrees C followed by centrifugation as described for 10 min at 18,000 g. SP:SAMPLEPREP_SUMMARY Supernatants were maintained at 4°C until analysis that same day. SP:PROCESSING_STORAGE_CONDITIONS 4℃ SP:EXTRACT_STORAGE -80℃ #CHROMATOGRAPHY CH:CHROMATOGRAPHY_SUMMARY Positive C18 CH:CHROMATOGRAPHY_TYPE Reversed phase CH:INSTRUMENT_NAME Thermo Vanquish CH:COLUMN_NAME Phenomenex Kinetex C18 (150 x 2.1mm,1.7um) CH:SOLVENT_A 100% water; 0.1% formic acid CH:SOLVENT_B 100% acetonitrile; 0.1% formic acid CH:FLOW_GRADIENT 0-0.5 min 5% B, 0.5-1.1 min 5-95% B, 1.1-2.75 min hold at 95% B, 2.75-3 min CH:FLOW_GRADIENT 95-5% B, 3-5 min hold at 5% B CH:FLOW_RATE 450 uL/min CH:COLUMN_TEMPERATURE 45 CH:SAMPLE_INJECTION 6 uL #ANALYSIS AN:ANALYSIS_TYPE MS #MS MS:INSTRUMENT_NAME Thermo Q Exactive Orbitrap MS:INSTRUMENT_TYPE Orbitrap MS:MS_TYPE ESI MS:ION_MODE POSITIVE MS:MS_COMMENTS Resolution 70,000, scan range 65-900 m/z, maximum injection time 200 ms, MS:MS_COMMENTS microscans 2, automatic gain control (AGC) 3 x 10^6 ions, source voltage 4.0 kV, MS:MS_COMMENTS capillary temperature 320 C, and sheath gas 45, auxiliary gas 15, and sweep gas MS:MS_COMMENTS 0 (all nitrogen). Data converted to mzXML using RawConverter. Metabolites were MS:MS_COMMENTS annotated and integrated using Maven in conjunction with the KEGG database. #MS_METABOLITE_DATA MS_METABOLITE_DATA:UNITS peak area MS_METABOLITE_DATA_START Samples CT WT 1 CT WT 2 CT WT 3 CT WT 4 WT 1 6h WT 2 6h WT 3 6h WT 4 6h WT 1 18h WT 2 18h WT3 18h WT 4 18h CT RagA GTP 1 CT RagA GTP 2 CT RagA GTP 3 CT RagA GTP 4 RagA GTP 1 6h RagA GTP 2 6h RagA GTP 3 6h RagA GTP 4 6h RagA GTP 1 18h RagA GTP 2 18h RagA GTP 3 18h RagA GTP 4 18h CT WT 1 + Conca CT WT 2 + Conca CT WT 3 + Conca CT WT 4 + Conca WT 1 6h + Conca WT 2 6h + Conca WT 3 6h + Conca WT 4 6h + Conca WT 1 18h + Conca WT 2 18h + Conca WT3 18h + Conca WT 4 18h + Conca WT 1 6h + Bafilo WT 2 6h + Bafilo WT 3 6h + Bafilo WT 4 6h + Bafilo WT 1 18h + Bafilo WT 2 18h + Bafilo WT3 18h + Bafilo WT 4 18h + Bafilo Factors time (h):0 | Dead E coli treatment:no | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:WT time (h):0 | Dead E coli treatment:no | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:WT time (h):0 | Dead E coli treatment:no | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:WT time (h):0 | Dead E coli treatment:no | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:WT time (h):6 | Dead E coli treatment:yes | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:WT time (h):6 | Dead E coli treatment:yes | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:WT time (h):6 | Dead E coli treatment:yes | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:WT time (h):6 | Dead E coli treatment:yes | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:WT time (h):18 | Dead E coli treatment:yes | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:WT time (h):18 | Dead E coli treatment:yes | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:WT time (h):18 | Dead E coli treatment:yes | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:WT time (h):18 | Dead E coli treatment:yes | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:WT time (h):0 | Dead E coli treatment:no | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:RagA GTP mutant time (h):0 | Dead E coli treatment:no | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:RagA GTP mutant time (h):0 | Dead E coli treatment:no | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:RagA GTP mutant time (h):0 | Dead E coli treatment:no | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:RagA GTP mutant time (h):6 | Dead E coli treatment:yes | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:RagA GTP mutant time (h):6 | Dead E coli treatment:yes | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:RagA GTP mutant time (h):6 | Dead E coli treatment:yes | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:RagA GTP mutant time (h):6 | Dead E coli treatment:yes | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:RagA GTP mutant time (h):18 | Dead E coli treatment:yes | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:RagA GTP mutant time (h):18 | Dead E coli treatment:yes | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:RagA GTP mutant time (h):18 | Dead E coli treatment:yes | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:RagA GTP mutant time (h):18 | Dead E coli treatment:yes | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:RagA GTP mutant time (h):0 | Dead E coli treatment:no | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:WT time (h):0 | Dead E coli treatment:no | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:WT time (h):0 | Dead E coli treatment:no | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:WT time (h):0 | Dead E coli treatment:no | inhibitor:none | Sample source:murine bone marrow derived macrophages | genotype:WT time (h):6 | Dead E coli treatment:yes | inhibitor:150 nM concanamycin A | Sample source:murine bone marrow derived macrophages | genotype:WT time (h):6 | Dead E coli treatment:yes | inhibitor:150 nM concanamycin A | Sample source:murine bone marrow derived macrophages | genotype:WT time (h):6 | Dead E coli treatment:yes | inhibitor:150 nM concanamycin A | Sample source:murine bone marrow derived macrophages | genotype:WT time (h):6 | Dead E coli treatment:yes | inhibitor:150 nM concanamycin A | Sample source:murine bone marrow derived macrophages | genotype:WT time (h):18 | Dead E coli treatment:yes | inhibitor:150 nM concanamycin A | Sample source:murine bone marrow derived macrophages | genotype:WT time (h):18 | Dead E coli treatment:yes | inhibitor:150 nM concanamycin A | Sample source:murine bone marrow derived macrophages | genotype:WT time (h):18 | Dead E coli treatment:yes | inhibitor:150 nM concanamycin A | Sample source:murine bone marrow derived macrophages | genotype:WT time (h):18 | Dead E coli treatment:yes | inhibitor:150 nM concanamycin A | Sample source:murine bone marrow derived macrophages | genotype:WT time (h):6 | Dead E coli treatment:yes | inhibitor:50 nM bafilomycin A1 | Sample source:murine bone marrow derived macrophages | genotype:WT time (h):6 | Dead E coli treatment:yes | inhibitor:50 nM bafilomycin A1 | Sample source:murine bone marrow derived macrophages | genotype:WT time (h):6 | Dead E coli treatment:yes | inhibitor:50 nM bafilomycin A1 | Sample source:murine bone marrow derived macrophages | genotype:WT time (h):6 | Dead E coli treatment:yes | inhibitor:50 nM bafilomycin A1 | Sample source:murine bone marrow derived macrophages | genotype:WT time (h):18 | Dead E coli treatment:yes | inhibitor:50 nM bafilomycin A1 | Sample source:murine bone marrow derived macrophages | genotype:WT time (h):18 | Dead E coli treatment:yes | inhibitor:50 nM bafilomycin A1 | Sample source:murine bone marrow derived macrophages | genotype:WT time (h):18 | Dead E coli treatment:yes | inhibitor:50 nM bafilomycin A1 | Sample source:murine bone marrow derived macrophages | genotype:WT time (h):18 | Dead E coli treatment:yes | inhibitor:50 nM bafilomycin A1 | Sample source:murine bone marrow derived macrophages | genotype:WT L-alanine 8388805 10821440 10659070 9481837 8927137 11527780 12397570 10415410 27528170 17921080 15796310 17727980 17446460 13142740 12182750 6237980 12578520 14539130 14035690 7603118 27749100 23600980 27661940 11547300 12647880 13086780 12833120 15278450 12932820 11324820 18012160 15793180 21424930 10060080 25337300 33462330 7914716 7790308 6211872 9519207 4450578 3151321 5035226 4907696 L-arginine 2730558 2838388 2621865 2138638 2504908 1539459 2259208 1524562 4182437 2432184 906706 1473723 3193514 2742951 5056814 1666932 1918251 3689634 2016978 1063429 2776749 6258934 4724733 1337032 618667 685522 732824 820460 638564 703368 897472 849436 1357975 108838 452078 1243267 775958 777122 725013 1575263 327585 199693 345442 248327 L-asparagine 904648 1049418 870329 1053401 1117268 919700 1435882 831383 3656331 2232398 1631009 1390939 1734270 1219610 2000925 651055 774500 1394084 1071612 446436 1882336 1806049 2712486 1558174 990136 859969 715891 732485 776322 796397 811765 651812 1072290 504898 922038 1119666 478077 472396 234470 569822 279676 144567 220522 267179 L-aspartate 2701537 4244160 2287852 3197569 4124442 3567604 4065193 2754584 3105240 2670231 2068126 1408054 5188996 4735642 5138405 4656068 2018305 3493532 4412594 2305174 6001300 3952874 4921258 3502369 5784886 8113554 5153232 5774552 5666642 6274220 6227442 5421108 4619240 2335654 4525170 6023278 6464788 7380610 4309642 3708447 2685322 2251818 4172430 4004788 L-cysteine 1478168 2418984 1313167 1942622 2697690 2563829 2527244 2116580 8927877 8100306 3402101 5698126 2881163 2252166 5599016 763859 3286227 3137909 4345658 1340579 5423376 4767522 10192780 5609120 1071379 1353870 871716 704995 941878 801623 1064818 828055 1533238 889115 1363563 1487792 490777 598924 296640 471356 155854 102232 205760 139865 L-glutamate 29077130 43319630 47861610 54992900 46114280 43382600 71956650 48886660 22507940 25577160 35667780 43352050 58450500 35531670 60421420 48935640 44323950 53428430 52564360 38411560 25155440 25502230 26615680 28230480 51664130 65632000 58355920 63735390 55800140 54422280 76988150 59197190 58943680 28132980 49405920 74804520 47012860 55323120 29887240 39546770 16384390 12622350 20119600 24342980 L-glutamine 6040888 8238820 6788126 7795670 4342649 4980248 6827034 4328003 14389640 8510538 9666094 7393368 10250400 7051714 16533760 4467110 3428345 6964068 6807946 2469100 6669086 5700436 14682070 7742622 3979316 5348408 4220154 5646516 3396229 2980389 4538924 3478920 6053608 2989103 4572272 9185137 2512492 2775719 1543237 3902081 1499991 756520 1594344 1623138 L-histidine 375280 438887 308743 399687 341366 305218 374138 389105 2156950 922572 444954 512924 709097 624734 881631 395474 348684 673898 610832 207570 1035667 1140681 1212630 598096 820717 703609 414149 612043 754913 475703 480665 562040 678345 293898 631130 803183 586623 480343 224298 458549 166977 181875 164539 184883 L-leucine 24417130 23470910 18146240 18789870 16658000 17080860 15905960 14424260 89163460 50202190 20810720 28195770 46568140 41684460 77131970 19335350 35272380 51331440 54200030 16400460 75630100 69022460 131195000 42797020 11027980 12121550 8812289 10451180 9574393 9558289 11641820 11437460 25160500 9739357 20138910 28756020 4871088 4771448 3471238 6652090 3300345 2033584 3630550 2432872 L-lysine 2021120 1916859 1252721 1654608 1587758 1206322 1358127 976947 8697425 3391946 1215086 1641286 2611839 2276711 4335272 1173608 1279102 3303299 3127888 650212 3975045 3871110 6979706 1894724 902573 1049574 681347 686646 917942 755868 775789 650869 1656625 597443 1162514 1200930 570626 488529 253791 444347 163241 115003 182284 218942 L-methionine 5982102 5338784 3894193 4414524 2129585 2102308 1893318 1986163 24888930 11541020 4155211 5199142 6409476 6064158 10897480 1960142 4685302 9618760 7771868 2032440 13367460 14837390 23760030 8271216 1330789 1538639 990540 1142598 1128354 1081539 1499216 1236774 2431504 1007956 1957952 2877406 572335 481283 368167 636705 226436 152306 317181 163394 L-phenylalanine 5593366 7144422 4050418 5024960 3251624 3339287 3337881 2966541 26965750 14409550 5765152 6758070 8260496 7696000 14014590 3604611 5534842 7762374 9451824 2641655 17363550 12808710 29819030 10929630 1797040 1760081 1137121 1480243 1342668 978966 1709109 1647657 3338741 1016161 2257973 3741027 856733 835081 466743 646280 399987 359887 484363 343534 L-proline 7451342 10292790 10459270 8635265 11388750 11357400 10058050 10088570 33306270 23139040 14172400 14780480 16377320 11980750 14296460 5057927 11487360 14163820 10388110 5176450 27285080 23925120 22798340 11544680 12370240 14341100 12317090 12459750 12698770 12436770 15907050 13174350 23836680 11575360 21444340 28383430 6761916 6235048 3969949 5825792 2640677 2300197 3525299 3013258 L-serine 1173211 1113204 741086 1027238 847587 819966 965115 798207 6546974 2762638 1872206 1449313 1695366 1564897 2589113 930985 616675 1500645 1153774 441064 2590293 3231025 3400743 2149624 1334725 1270916 774771 1114212 957620 1099525 1229625 973367 1523454 866307 1094265 1490944 1639011 940133 448990 879968 615204 579501 446370 604877 L-threonine 1721597 2318852 1640288 1623705 1452557 1633734 1674958 1304654 9899872 4007473 2109702 2734410 3241790 2862448 3660783 1528492 1595309 2838622 2810375 783204 4714218 4931188 7005594 2816099 2136436 2471446 1561942 1898393 2086578 1993799 2124157 1800199 2732534 1299861 2161182 2614334 1608556 1422293 783257 1311468 535755 456006 490110 543028 L-tryptophan 643603 891406 596244 562848 647073 815446 495124 722612 4179879 1900657 754789 955405 1439813 1509330 2174187 513589 1249829 1386768 1669571 500935 2125124 2474162 4140086 1624850 652658 664415 537120 472649 439995 367591 517168 654468 723781 366236 837831 951952 325863 232534 212032 354166 141666 94178 124071 85026 L-tyrosine 2871604 2661596 1989125 2250860 2057642 1931627 1863808 1695330 20841780 8771151 3426111 3150794 6274454 6279642 11247050 2978530 4321282 6580084 6937320 1908761 11894110 11930340 20696220 8943396 1862948 1898216 1060742 1566342 1406618 1195070 1837237 1690561 3501499 1568525 2802353 4189071 831202 846079 508371 952048 470627 339217 442023 364015 L-valine 10446480 11068210 9027383 9324004 10373750 9793396 11083230 9991813 42880510 25868760 16872580 16530150 21848260 19354850 31562360 13029700 16378700 22683700 23032930 9629896 34007840 33220480 53565950 27749130 9116035 10126520 7375576 8089988 10091940 9939913 11575290 11528120 18804120 8713024 17420600 26036720 8015454 7946710 5852984 6843438 4713804 4382427 5187344 4936764 AMP 4573732 7095966 6204500 6656660 5793410 5460672 5353518 5135002 6159712 8523457 6240248 9289069 20186360 13622930 18435880 6254784 9569828 10332230 8987360 6701820 15593830 16703310 13859900 6652454 8174680 9225091 6564100 6627060 3959961 3921187 5216126 3913357 3820716 3160561 6255334 6849632 4219448 5271182 1741123 5186148 932684 593033 994427 1011472 dAMP 14708 8763 19896 8360 162077 89546 67159 135219 53350 80670 66923 81882 27796 22147 10862 31706 148316 126486 92794 166370 82935 88118 77314 67166 26143 19257 36254 36221 51907 51923 45481 48095 52970 22265 57373 45748 30556 18662 74215 38529 5321 6854 5578 4172 Adenosine 8148984 7988904 5004394 5168028 2392860 2175455 1333747 1241974 1820480 3333335 2318642 2485852 3473717 3115096 4611098 1204694 2591379 10310830 4295098 734057 2540710 3319314 4511383 3605792 1261468 1590209 619302 770360 945660 557524 1035107 608371 888239 683759 1287566 1198055 1210802 663351 166607 657391 287207 168830 226071 286417 Adenine 1271713 1446262 891372 967788 926600 949916 691053 655083 776557 647820 738244 892685 1227638 996705 1236707 655395 892701 1954420 1254564 523585 1121985 1325206 1584207 881220 643868 749034 453351 560212 617133 612386 839691 602228 857049 614859 878353 846940 835008 641211 315403 546918 328253 277634 306548 288745 GMP 279081 794230 351288 507174 238497 158247 246278 186856 1737522 1648315 1038923 1221580 1024929 627718 685530 213587 496262 641893 341927 344573 1658255 2234545 1718856 870399 313421 497520 344396 225368 361836 277055 547329 239903 815187 264900 992966 1082117 226434 211854 80024 267272 49477 37064 47895 42324 Guanine 203888 193856 128872 132605 48389 88801 41622 72084 690820 387682 121949 134836 268774 223645 300371 68981 191020 321771 269291 63879 459664 437223 782430 289453 43476 61793 93535 93359 117891 122823 126801 117824 67196 38175 71696 95763 49499 64832 46927 41063 29800 29291 49266 77610 CMP 284014 381210 379786 366535 232022 279186 271265 197975 378753 426804 291940 399330 495517 372841 430591 234230 225526 242610 204125 151394 297106 397997 485691 396535 387749 528113 395850 345224 206903 188570 273549 225226 220522 147460 369511 387639 137842 139633 92174 151385 24678 18997 27054 17168 Cytidine 174953 417788 226164 242097 71269 114653 111701 73311 1665474 1512862 625713 691961 264240 147853 294816 222003 124477 264991 94222 58030 1042655 996436 966742 707959 686371 806579 404346 393082 92502 29649 51930 44516 144715 50454 171632 121118 33529 30449 33226 20299 11382 4627 20238 4580 Inosine 1452131 2433946 859586 1913649 343469 507091 423831 491363 4548018 3822259 1733183 2428965 674625 457245 941474 332431 680034 1565520 862758 89490 2867553 3527761 6671234 2734454 452566 558210 274683 416634 260213 232905 412984 207302 565047 621344 1077804 911116 261258 100127 33996 248289 43778 14811 30609 46433 Hypoxanthine 12446560 18840650 7450388 15538130 4495704 5906536 6028152 5221774 52544760 43763460 21216100 27860580 6703046 5250148 8638140 3357483 6666448 13153850 10374830 2013713 39508900 42146730 69979740 24035940 6238524 6720398 4285132 5104690 5135804 4278652 7847188 4854950 14495410 11063600 18233800 16454220 2887508 1902126 915775 3342623 715660 555391 827497 963247 Allantoate 86180 87884 65626 67571 271768 149437 155355 135138 589695 308416 145924 241092 123350 110653 67717 59726 111383 150388 108367 88246 454164 537304 528654 267947 192498 148415 104378 129001 266992 161510 219618 168934 1300678 663807 1115713 704438 152226 100206 58540 92577 168795 122437 155810 182101 5-Hydroxyisourate 112427 152330 174634 121073 73779 71168 104362 105575 227330 200945 155567 153740 389398 169331 359172 77078 125706 166379 90260 45452 220027 102260 301160 170489 95443 192601 109880 180917 93149 51637 140274 63560 122167 12670 153103 276680 36315 77385 20688 85393 6343 3185 5408 14977 2',3'-Cyclic CMP 0 6332 0 0 35260 10662 32083 18420 14918 16914 16984 57099 14148 0 5445 0 42120 45637 33101 32893 3702 17017 35608 35792 0 15385 0 0 9644 8316 9230 5809 14887 4303 20011 21175 18732 9330 9422 17673 7475 6501 6786 4053 5-6-Dihydrothymine 171951 207512 208725 232775 190943 232829 236866 233211 607325 592051 382016 685901 472732 400604 293141 246257 356369 433519 505283 279797 1229724 1062720 1500165 603894 287643 361177 284853 319106 303133 261159 301184 334058 1167003 398663 1376340 1541199 234807 307105 206071 228884 239040 168028 261335 274440 Pyridoxal 155979 122810 73891 76757 122205 159525 93109 108408 231068 153555 164460 185513 92127 63614 179096 84456 70125 121115 147697 56359 175010 98418 225990 64745 75590 67959 44836 74002 91530 57488 91199 73564 94997 50333 91017 118364 61693 47132 34950 63539 35461 36439 24713 33727 Nicotinamide 8040062 13978040 5296324 9778779 6831882 8879872 4156962 7236104 35434370 50825060 23717560 37715940 13771140 12549830 17985190 5720348 12016700 16980930 17217900 6922462 32854100 38224400 49761920 14857480 7592060 15177880 5516910 6232252 7740494 6003742 4839948 5188092 12742580 5582986 16501040 21176190 2458379 2167453 1054944 2235846 638112 660528 755639 665907 NAD+ 205263 269378 373915 417278 247906 334092 321650 421382 39051 44617 36949 59834 1061681 707784 1035312 724960 520782 618938 426095 354106 73432 49622 141747 14117 417576 519936 530870 615997 261405 157100 439785 339433 19403 4028 118774 108308 156180 164137 158001 234456 37750 26346 61353 28182 Glutathione 15055960 19428380 14434540 17049780 16232500 16069090 15280180 14280020 22812800 30228160 21398950 30018840 32715950 25797330 37135660 23430780 19669300 24132660 22635190 18060190 32481350 31089090 35584900 20456970 22557420 24422960 18672310 23821000 14315110 13047290 16426310 13913240 19996130 8167302 19960290 33521570 9877283 13755600 10554450 11557270 3315045 2477409 4586681 3188627 Glutathione disulfide 27567 17729 29696 5876 14614 15859 20983 23751 53604 67804 100529 77015 40154 20669 17093 52491 17816 17287 6761 22333 133279 63768 58488 62946 76375 103170 114715 176630 117160 60273 91679 43875 115202 55190 93448 121598 77913 89209 59335 90858 31640 15795 33477 29098 5-Oxoproline 12107150 16490500 18932780 20137990 15558940 14913680 24187370 16405980 15143160 12558690 16316840 16953630 23061790 13583510 26772670 16383080 13116760 17868660 18966190 12464190 11272920 9532512 14568260 11271330 17363060 21748390 19335060 21169350 17612590 17232740 24274980 18290530 19129700 9600949 16421990 24643920 14786230 17754570 8646869 13805960 5729792 3977751 6460334 7512650 S-Glutathionyl-L-cysteine 125116 116298 116233 116990 63173 63568 53041 57064 75693 113978 236834 112683 77866 107391 55234 124968 20391 89512 54834 38117 77694 264508 100328 173600 199233 144796 126303 139595 121977 100862 106010 89559 483691 245937 632754 537350 58831 94627 44353 53829 61654 11409 78085 58520 Cys-Gly 180100 222758 116451 141615 164905 215008 142631 121989 217237 251078 178940 212869 300565 327104 424652 171985 196531 389918 291746 116481 255423 340442 486989 146912 144127 176131 114726 149644 101259 70644 119304 96886 124808 57199 136651 233925 87551 103056 67774 61313 22805 22166 26702 23980 gamma-Glutamyl-Se-methylselenocysteine 47017 31544 30828 28091 105768 60885 77515 47944 298133 259757 126364 157590 36602 34093 30916 9142 20784 67810 61143 21098 186952 178623 204390 114818 76690 33122 38126 54696 120600 76791 90969 78355 478160 256695 375863 256812 68539 41669 22231 28710 88356 66671 83248 78758 gamma-L-Glutamyl-D-alanine 418978 542391 291875 478359 436626 554815 396186 367403 554573 632614 385412 564069 750860 722014 1125957 295453 503572 1216125 982565 347665 513718 977219 984242 400615 136217 130794 104830 132278 121105 82713 110047 117235 198845 51171 179806 258236 102499 104580 71050 79354 36312 44722 76775 37852 gamma-Glutamyl-gamma-aminobutyrate 610769 723187 463599 415855 663783 737963 523121 524044 920309 827723 379968 473988 1172777 1260177 2015552 504057 818474 1267777 1579476 501370 690447 1131472 1946866 652482 96647 116558 70654 68590 66467 50982 74051 56170 47200 14330 30300 60983 39813 41236 20918 24355 0 0 4588 8674 gamma-L-Glutamylputrescine 428252 300113 191524 336447 249296 203531 156555 210297 297442 172806 47772 143887 511529 462701 641984 112792 383543 671122 620030 171102 148747 291279 720542 119992 2037 0 0 1759 0 6940 2286 3837 0 0 0 3226 535 2243 0 21902 3131 0 0 2828 (5-L-Glutamyl)-L-glutamine 157160 199365 125306 145011 164766 185700 165311 153506 180747 198157 125368 182511 264794 340013 450243 115570 129749 422739 392411 106098 145602 307112 436268 126188 85086 68543 57921 93251 59880 42331 98423 77642 109194 59216 116959 158954 70307 52114 35219 50104 29412 32830 31569 32344 L-Homocysteine 35252 17893 33026 7568 31275 35066 51881 33393 67758 32000 6231 19015 69116 47211 70456 21750 40512 72111 61108 22353 61929 73095 102292 33697 23318 14828 35291 20660 37307 18310 23368 26197 0 9032 1779 24256 0 6043 0 0 0 0 0 0 Cystathionine 65838 107150 49918 74294 103865 106547 68266 65624 51127 85040 33559 68049 86868 118365 256620 18861 111850 271875 202839 41090 34420 123834 162112 27983 30198 24507 7664 5140 1463 0 7012 7927 23653 7537 18563 32075 5489 0 5382 7134 3525 0 6728 5546 Dimethylglycine 848738 1357672 945121 1141119 14945020 7859634 12625150 9477931 4293250 3343201 3997074 6123040 2027271 1212820 829907 548394 8951067 11295640 6550606 6310338 8394184 6577930 8510986 3265058 1484679 1496332 1297568 2214519 1006290 908183 1228051 1337562 1500199 681668 1993941 3056451 890092 1119428 798558 1105639 414698 293438 497480 473387 Phosphoserine 0 78994 146104 80930 65378 60264 136030 131169 55477 71451 71260 40285 192410 62501 88578 102053 148825 161158 96998 66525 49141 45277 21938 80727 137602 231197 184089 240797 199625 97835 227983 126813 100107 61604 103459 191429 88153 127699 73433 158623 63749 87687 81624 133810 S-Adenosyl-L-methionine 17401 68943 24704 47554 50124 58168 34311 53677 180225 141995 73015 123194 72320 45678 91116 31697 60634 92056 86448 30662 260892 372575 204804 64164 58480 20689 42156 49582 50112 28853 58805 19930 61518 38445 91041 87515 42756 47700 12524 50893 11082 10054 20769 30548 L-Homoserine 1721597 2318852 1640288 1623705 1452557 1633734 1674958 1304654 9899872 4007473 2109702 2734410 3241790 2862448 3660783 1528492 1595309 2838622 2810375 783204 4714218 4931188 7005594 2816099 2136436 2471446 1561942 1898393 2086578 1993799 2124157 1800199 2732534 1299861 2161182 2614334 1608556 1422293 783257 1311468 535755 456006 490110 543028 Ornithine 160049 149208 139103 120119 462991 264844 284811 146007 1687498 684108 322516 591515 233082 212597 194735 86817 154864 261690 282646 164350 864466 1084450 1373698 422363 362353 281026 201703 224320 485436 283925 421388 293033 2623759 1015400 2003018 1610676 280688 191220 80039 144627 192942 169821 220716 306949 L-Citrulline 1891744 2100952 1695789 1852795 5854706 3412461 3994882 2605518 18622170 12478440 11521970 8865239 7081442 4251057 6454868 3618261 4136036 7821962 5869730 2126268 10185670 11626260 12081350 12353690 4656500 6289862 4024379 4338868 7954910 6699754 8220668 4313544 53133490 13751030 37560570 28410570 4254824 4486198 2378766 4232124 2118050 1388144 2014252 2469656 Argininosuccinate 81040 115924 97567 85527 308370 168914 247718 73176 93538 164048 413366 126603 116149 109649 132554 73840 131142 184761 84653 54066 103551 143484 112050 238170 62994 130170 103521 77641 82028 121779 147559 88970 167380 55107 186488 245898 49759 68371 45426 79950 14782 23675 25362 42876 Cadaverine 9812 0 0 0 144617 104954 96112 100098 255452 372734 72236 207447 0 0 0 6311 87410 123823 74589 113138 155946 303858 458316 120415 1633 5261 0 0 323324 460811 407494 338336 1451108 654830 1082447 656683 223825 297223 418164 259197 273368 305292 301767 214723 Putrescine 138902 272371 147814 157646 557559 827090 520162 806604 1102757 1468574 430945 1005139 386477 250819 400402 357942 1355096 843679 705028 1004104 1671228 1613671 1892394 1049988 236482 500595 209884 333965 640202 1192006 905113 947304 1431942 1048277 1415704 1239606 723952 880782 644554 793830 266029 185512 232971 200455 Spermidine 13030600 14601720 12786280 15359020 10656950 16608410 10134350 10617580 16326230 17221780 12588180 11534750 10170300 11171260 18130860 9481277 14643620 8958908 12709670 10223740 12069380 12122240 15908210 9483373 14357920 13000720 8979482 10051410 4333640 4102470 3084286 3584595 3501604 5852960 4979336 7180130 2146663 4170674 3773150 1761363 1048060 524072 1392602 945324 Spermine 1578057 665174 1890067 1407252 356685 475205 354326 452834 218046 160692 223682 81848 247624 393341 653594 313994 330996 315641 372183 326318 114962 240176 262732 376301 869465 371460 754383 529283 141557 75624 367652 367598 114692 161112 250261 159056 142992 108432 136125 181797 86307 31130 114968 58360 Carnosine 61019 62408 48804 59815 55122 38327 49835 47969 108099 86770 37378 83079 115818 95470 148301 23903 67543 114643 129947 26133 111166 72590 217579 63759 9253 27239 12143 29708 14585 0 15331 5094 6357 0 6184 7656 4994 5905 5860 11496 0 0 0 0 5-Guanidino-2-oxopentanoate 2618 2349 4536 2774 29199 5979 12971 4466 13637 16545 13001 2677 2341 2141 2476 5178 11370 11633 39438 23140 16719 52039 16813 13857 2396 9136 3115 2162 69108 128277 104682 85574 415518 74922 187972 264195 108876 118975 87474 47339 17333 21874 17368 27631 Phosphocreatine 965326 492349 346088 548709 205755 592367 182574 578625 1289876 1436925 875182 1617443 1172484 1300649 815686 1116239 1145289 2356543 2404188 650222 982502 1314492 2128617 1097320 472495 717472 229769 908847 279349 131997 121400 300647 5207592 4135629 4228976 6355802 271486 293293 286608 281025 421457 286039 385946 316246 Creatine 19657720 37657610 25650200 35613380 30858090 33692180 30970460 33203660 128235500 146369300 111067500 186778200 67265470 71246860 73802100 46868760 54556910 61870180 71204230 44747540 145510400 233251100 247759000 129814900 49305190 59135450 42064480 59630960 43406060 33061700 36632230 49118860 306108500 100415300 401655300 453405700 46628930 51207720 39333980 36085330 39177800 28825410 45523310 40798910 Creatinine 833905 1063924 1066594 884436 1162008 1181450 1107365 1243754 2129313 2171232 1506122 2090380 1893469 1412809 1474917 973777 1791194 1719498 1532145 987590 2659923 2692291 3570027 1912134 1293665 1437541 1188058 1587405 1368731 1050066 1347016 1375594 1784972 1059574 2388677 3121100 885384 994821 1299251 1367523 693307 355582 583474 565188 4-Acetamidobutanoate 68957 75074 66642 58536 63167 63051 61331 53174 82982 68969 61447 75439 99548 117159 101683 39283 57733 92672 86039 33182 60794 70214 99390 34415 60877 62786 49850 73695 54871 34172 61898 60500 53271 46679 52896 57428 34537 44644 62371 49191 39949 38649 46026 43088 N-Acetyl-L-citrulline 345658 393797 238209 322760 182606 190233 165705 152678 377287 282661 92203 166908 344601 373306 566995 94610 274264 606317 586411 97553 210017 440146 637488 175200 0 13006 12925 5733 8117 7811 7901 7085 13138 2789 1960 3050 13288 11736 17195 6156 8261 2818 13391 4094 N-Acetylornithine 1434420 1553091 925232 805657 787062 937509 668904 699526 1514386 904783 517186 645506 1487239 1787421 2771344 469003 1284044 2709891 2482929 522118 930242 1735462 3679787 944661 142778 135523 82107 104909 98444 78913 150084 93377 223429 177370 275476 344554 82390 75365 31311 76507 56380 63343 82686 61460 Guanidinoacetate 1553310 2718378 2785209 2782319 1267392 2372044 2733730 2561652 1331601 1679221 3417603 2463550 6383238 4671890 5877532 2914669 3115061 4010464 4682666 2272795 3497954 2542373 4201784 1380783 1923211 2747928 3531165 3405359 1648357 1573867 3629500 2800851 1080522 495998 2630722 3107453 473199 601898 391606 522648 123873 75285 171850 146576 5-Aminopentanoate 10446480 11068210 9027383 9324004 10373750 9793396 11083230 9991813 42880510 25868760 16872580 16530150 21848260 19354850 31562360 13029700 16378700 22683700 23032930 9629896 34007840 33220480 53565950 27749130 9116035 10126520 7375576 8089988 10091940 9939913 11575290 11528120 18804120 8713024 17420600 26036720 8015454 7946710 5852984 6843438 4713804 4382427 5187344 4936764 Pantetheine 50817 43142 18397 42726 21151 39178 7913 18772 24208 24205 0 15075 43836 69016 63174 18347 39382 108592 75204 7604 13653 17613 134489 51846 0 0 0 0 0 0 0 0 0 6588 0 0 0 0 0 0 0 0 0 0 Pantetheine 4--phosphate 67932 42560 39659 44354 60930 106364 65746 66182 76035 80593 15992 57704 84536 72842 146174 47949 97947 131381 134299 50437 71965 91597 158491 23259 77051 33646 14176 40606 39521 16445 5009 15109 19585 3687 19613 16296 3673 23547 25819 1534 0 623 0 4711 Hypotaurine 1354273 1732965 1455278 2037817 1318892 1499910 1718793 1341650 2066643 3080737 2342971 2916090 3916340 2875577 2911742 1910751 1615220 2822247 1793363 979420 3055917 2626304 2348462 1661197 2310636 3087801 2034573 2992208 1949265 1552516 2123670 1698041 1872853 951812 2956154 3606908 1854741 2045068 917852 1536771 835722 562740 917152 1292645 L-Methionine S-oxide 61384 56212 48507 46777 93019 91896 84156 66483 177835 109432 71154 99645 177943 136549 181742 138341 89522 131411 137272 39389 163487 166670 223949 157754 79708 84841 58443 54261 73452 83008 89332 71197 174628 110605 177576 206839 68205 43629 22569 47921 23955 20196 13235 15461 5-Hydroxyindoleacetate 0 0 802437 648465 1129763 0 0 573790 0 747296 0 442402 0 0 556271 503375 1458000 267310 1862944 677558 950314 0 2091934 669604 887613 589004 0 1232082 1055978 0 563025 278621 342181 0 1107164 0 350120 538393 0 663763 169575 181631 292373 124210 3-Methyldioxyindole 37689 43694 41898 44221 38520 54314 32383 48566 172137 99461 61609 56168 74339 78269 100523 34970 68905 70503 86184 33616 104828 107836 175078 74418 36796 37361 41710 34380 29219 28826 34871 38811 43818 34942 49989 52105 22812 25226 24417 31509 21911 25826 39610 24129 Indole 66265 68490 46655 76997 51017 40140 42394 56746 314197 148711 59411 103157 76563 73921 146670 70830 57304 82721 130263 37015 154537 94205 351806 110390 41881 41046 26496 25591 38922 33564 21658 29833 43439 23306 22036 68731 24346 24550 20926 18991 19813 22003 24722 28205 Indole-3-acetaldehyde 44053 36095 8624 21550 27908 19636 18991 1924 60149 55320 49167 44933 37347 65230 35558 16911 37895 51372 49307 1814 72740 22146 61417 76772 2921 4109 1952 3502 26133 19010 30517 7284 185159 52376 132336 69114 7762 5695 6341 16822 3989 4859 4209 4323 Indole-3-acetate 186316 261121 293074 177652 184737 168950 193444 166167 260695 161049 124886 201453 288437 179415 485933 245555 240763 162941 157681 99452 470146 296665 436586 153020 280442 265618 330618 320099 134499 74927 90481 235765 67953 81319 104403 144284 128208 110003 130745 191136 86526 36265 71635 45606 Indolepyruvate 53516 33238 43238 35353 32774 26787 17201 52787 163506 90801 46322 30126 52846 79545 131100 37707 98367 143485 65513 56026 159565 162143 101157 100303 20611 8631 28969 25757 47623 25723 22034 37783 60582 34005 19334 55409 7041 21337 26729 16372 5886 6830 4781 3191 kynurenine 24694 24493 13392 14218 8116 20167 4089 25815 0 31755 8948 11716 11530 19542 31168 6573 36064 57994 48661 24605 17040 20579 17020 2466 17868 2399 6900 12123 6546 7086 2244 6232 13785 11619 11335 16452 12211 5633 10494 3031 8861 6318 4919 8825 N-formyl kynurenine 210634 242791 142556 205019 213880 213382 178792 171440 218962 221732 101955 125817 249227 328578 514756 104815 253403 497760 532625 109698 143654 186681 441038 69636 36907 20536 22005 14116 11245 23176 29248 11169 13017 27967 3716 33969 26601 20556 7643 11924 4874 3158 5751 8369 Anthranilate 192969 225583 347774 173451 244894 251652 1082047 305368 808422 493453 281333 396360 232798 246178 244232 106453 474392 296285 253053 173360 778769 571983 511548 304174 237259 299017 265316 250216 321506 195457 213134 326381 339174 127323 311289 323967 113617 110419 133425 98712 91715 65697 87668 69006 Picolinic acid 2361627 4737989 2484112 3782658 3864340 4916821 4960205 2760273 9138543 7722552 5668882 6730444 5589876 5264800 5401505 3628098 3749568 2204342 5449444 3173614 8726556 6587232 9619095 2945747 3734925 3938830 4500480 3202755 2830480 4003522 4307046 3311013 4046785 2963088 4581396 5114036 2555497 2893388 1003094 1431098 473186 664374 1099618 760713 g-Oxalo-crotonate 980551 1011319 883487 1108029 496896 524901 607031 823098 3046236 2962167 1858838 1902561 438062 424405 387074 225211 1116356 834942 564684 358770 4212404 2705233 3620509 1496161 592429 627958 586914 570857 741862 413627 867440 620698 1267305 728797 1461118 888526 231680 157975 152005 339699 61607 62151 88489 58230 2-Aminomuconate 16786 29316 28511 19485 18260 17043 20379 20393 29743 24053 15938 23876 22595 26190 51372 29299 30937 18115 15263 3712 59294 32234 40030 19757 38885 42573 34116 35962 10050 0 7961 21358 7792 4514 12662 21392 14505 3892 15214 17554 5351 0 5401 3581 L-Adrenaline 372850 392747 330235 396284 128897 165412 175139 181518 285444 252635 316698 287311 447074 468109 520946 373751 339734 258230 381061 276561 403219 421152 324187 144057 278614 250332 240347 297004 190807 264267 340491 356686 268096 189766 352940 309596 230971 171141 178993 134518 57212 22021 67066 54501 Dopamine 64794 47787 49655 18491 35770 28583 9601 40719 158405 84036 27572 43056 59235 79546 116687 9516 111482 151895 137606 38900 167499 159828 191416 87408 10363 0 19678 23586 22357 9568 19906 28736 59172 0 15601 15528 16534 8980 19724 18623 23228 15807 17067 16100 Serotonin 12665 16759 10793 15667 42984 22643 24261 19968 167720 123766 104760 55126 57521 47556 45458 24513 38461 57934 40779 17846 90925 77838 117526 97591 40750 42140 24558 47751 61661 44572 57370 44843 483719 118943 276442 264406 46075 47022 19856 33865 17360 5406 18671 27388 Acetylcholine 2087065 2646918 2636060 2539066 1689763 1909333 1723514 2578755 865556 837566 608129 1183031 4367664 4808858 4432238 1357631 2854431 2979915 2552589 1143556 1063073 922485 2774995 627701 1940533 2373007 1884621 2646468 1727058 1195162 1300296 1838343 726608 491335 849037 1457663 1185424 1257313 1033959 1040608 273492 269508 292655 222477 acetyl-carnitine (acyl-C2) 2135438 1636020 2983373 2715783 3876849 4003563 4127935 5227362 4436736 6235156 4911474 8505363 4949876 3660221 4282454 2774347 5684316 4847290 4229576 2992806 9300791 9269833 13041400 4628198 6205314 6357786 5563978 8851679 4018553 1604866 3908514 3288418 2859274 853167 4703394 5473094 2010219 2929969 1680324 2664706 373129 258432 609970 316872 butanoyl-l-carnitine (acyl-C4) 34837 45494 17386 22121 17464 11203 12046 20732 16461 44699 12302 23005 213708 67374 48615 56495 70172 36517 19928 18973 51166 50687 0 31097 35342 11308 17409 36013 14803 7988 13165 10225 13631 8460 8063 35439 19905 22744 13901 20501 11662 10016 9404 16265 acyl-C4-OH (Hydroxybutyrylcarnitine) 55439 27783 84260 47270 242821 182235 252825 216256 80390 37049 121392 59639 109056 91109 102037 79149 163607 209082 100365 82615 60954 89310 141639 120623 297847 263490 259373 384474 262604 107708 262824 228317 150693 39316 183785 281461 89380 150608 90340 102461 11823 15848 19230 26680 acyl-C5-OH (hydroxyisovalerylcarnitine) 43629 32349 44392 50144 42093 45648 35270 20973 44044 40940 35439 72048 82332 62326 112077 53468 48282 80478 69958 26412 58224 46293 106331 23942 25355 47666 19630 69713 24519 15807 28718 35570 29722 5084 53033 65279 11319 26129 15145 10856 4931 10477 5664 13839 O-dodecanoyl-carnitine (acyl-C12) 13505 21069 11962 7773 8033 18797 10762 5208 9459 21032 3194 3919 96681 42083 30733 48845 31957 19812 21026 18383 11699 11464 5052 17320 8870 12383 11998 10996 5438 9912 5777 4707 4218 3509 9074 9901 7628 3847 5815 10719 8336 2956 3679 5066 O-tetradecanoyl-L-carnitine (acyl-C14) 82369 97968 69514 59576 33596 68544 40656 39434 43542 76205 26954 60651 381913 212469 176139 142704 149645 107595 124569 77069 49440 128717 120494 49155 53644 62058 67482 70865 40639 35172 34050 39531 15178 4303 42576 43241 34863 31261 34304 27248 15339 5933 10519 12745 Tetradecenoyl Carnitine (acyl-C14:1) 17094 28678 7218 4509 8530 16220 2840 6261 2783 12139 2304 2854 136144 73814 53150 77351 42854 28152 55887 20645 15799 18664 36299 17295 10585 10997 2816 14222 12655 12392 3817 3173 3123 3086 1672 0 8047 4841 5541 8801 2734 2844 1417 4374 L-Palmitoylcarnitine (acyl-C16) 309459 422193 225011 365675 196509 362582 240885 150407 196138 366280 159758 282961 764619 711551 763374 322395 239143 447833 435840 233343 248687 351545 673526 191672 319905 449119 377442 301899 191010 186098 280136 284173 121119 96635 220787 280385 225463 240031 214373 100779 64966 95232 121398 90951 Hexadecenoyl-carnitine (acyl-C16:1) 48933 64138 40302 52314 56159 76043 54858 60683 35215 36746 20934 37145 284370 260301 208656 120084 151729 139001 156526 82102 40306 74758 113147 41652 53665 68717 60065 75578 38290 48931 51016 65083 22176 9228 20684 39433 39720 51921 39011 39295 28591 26818 29406 22020 O-octadecenoyl-L-carnitine (acyl-C18:1) 143213 217175 96694 150269 119557 160661 96290 79956 60245 85126 44548 102406 433370 455855 608638 198451 161050 323410 370487 111424 123682 130869 313205 67450 125354 144867 140342 159206 85690 106829 125319 153242 34183 21193 52974 79210 95872 72073 97845 29149 33346 46331 48220 36030 acyl-C18:2 (Linoleoyl-CoA) 14188 10642 9915 7800 9011 9353 8927 10607 1729 0 0 1083 28859 44490 30658 19025 19736 21718 22385 11518 2348 1151 1991 0 9381 9082 10110 7398 4389 7215 6203 8935 459 0 1199 0 5430 2162 2786 2196 1141 0 2506 0 Alanine 13C3 0 0 0 0 414472 230516 322415 225594 81779 48395 47957 23341 0 6236 0 0 118052 160772 155325 117872 28542 35639 89433 56492 0 0 0 0 329312 502399 566332 379133 303187 76065 243297 266195 396744 401227 338946 335684 48759 47204 48203 47549 Arginine 13C6 0 0 0 0 112967 19205 81988 48366 9279 0 0 0 0 0 0 0 19064 25130 53096 34762 0 9959 11194 9658 0 0 0 0 36746 61236 51009 39574 10746 0 6795 11190 25370 30572 17481 28349 0 0 0 6338 Asparagine 13C5 7311 0 6414 4013 296332 95103 145684 87705 33238 16299 19582 12388 3515 7326 7233 7028 27968 56919 57026 51250 11408 13182 14943 26300 6619 7701 9411 21313 288052 421944 387098 306578 306516 112222 206625 221754 445588 520579 392925 317507 54934 86362 83893 97735 Cysteine 13C3 2973432 5484074 3045842 4507438 4410765 5541639 5924400 3363013 10576470 9079226 6681022 8761923 6799182 6067228 6649060 4299590 4505238 2572949 6336978 3709007 9214987 7710990 11299810 3516151 4425558 4691462 5275668 3828193 3210180 4756452 5103596 3901055 4725742 3448535 5443392 5944642 2878043 3433256 1281598 1896997 621616 821268 1356885 903432 Glutamate 13C5 7111 8165 6870 3833 1343546 604473 1197837 558013 59120 85665 135632 169284 4929 6006 4588 4516 222467 297167 248940 350550 68549 66354 83459 110730 5691 6886 7175 7783 1709605 3120184 2455490 1430761 2590366 778395 1580590 2102249 1845351 2258407 1416477 1744796 270685 300748 284750 410580 Histidine 13C6 54773 60918 42460 49756 61405 39172 44315 45487 53132 38203 13400 26466 85454 89532 139656 24719 61033 143427 106642 22822 40810 57837 75945 38736 0 1486 0 0 96430 144166 104728 83813 89623 13619 37247 106348 88783 90953 54854 80636 7178 7204 4731 15310 Leucine/isoleucine 13C6 0 0 0 0 2914179 1225082 1569938 983150 550110 291112 185138 304795 0 0 0 0 1188387 1681399 2344032 1248122 418959 420119 733824 525566 0 0 0 0 703344 956475 1233646 808671 1169656 204341 557841 646047 268375 371647 253086 292851 32854 39835 36425 44194 Lysine 13C6 0 0 0 0 151287 79575 83764 66802 34993 29619 17133 36994 0 0 0 0 38574 69247 76368 35410 23096 24725 35448 24396 0 0 0 0 57006 87837 79236 59375 36987 0 23555 45914 40337 23597 16223 39390 4482 0 0 0 Methionine 13C5 0 0 0 8221 334986 135448 158337 110426 134330 98381 74456 105930 10502 0 12197 0 145737 178136 248345 147871 163207 186827 179376 46655 0 6830 0 0 106480 226992 168700 156634 114693 39579 95859 76866 57494 60628 32032 40207 6140 0 0 6492 Proline 13C5 4124 0 0 7465 1028731 607756 687386 498139 209544 163471 124240 159524 0 0 0 0 270343 373605 266102 246800 158867 150282 104400 127862 0 3944 6043 4322 557456 947137 1127597 691476 583648 168033 356800 469657 394600 556407 372925 348170 48770 60221 50963 37506 Serine 13C3 0 0 0 0 62123 22997 48991 24969 16961 0 0 0 0 0 0 0 8257 10600 29099 5616 0 0 0 0 0 0 0 0 74134 117554 89022 70877 89065 8828 67702 53526 84766 103774 53842 78699 9246 15312 5282 20848 Tryptophan 13C11 15689 16320 21086 21193 118292 62764 58482 36216 24794 22705 19810 24365 20414 15394 20017 26321 50358 49265 58467 34120 22100 23483 32257 22797 28761 17705 20251 21642 22674 35910 37352 30601 24392 26354 24269 16977 29392 24409 24309 15748 24782 27970 26233 16898 Hypoxanthine 13C5 16181 8465 10302 5864 1809636 1214130 1602060 1177491 6603964 5667306 4909314 5546089 17103 4331 8902 5976 920036 1984714 1267624 461586 4109505 4744350 6478172 4803073 8338 7230 10676 7496 761187 1093157 1487221 981316 2208994 688169 1765994 1825831 538624 451160 244583 764489 79941 81581 72463 99305 Xanthine 13C5 5584 1491 2011 2137 153004 59982 143850 26811 84001 81630 72969 96638 2730 6358 10652 0 26851 70102 63961 34280 23586 28162 39226 73653 4233 2249 2591 1852 163871 436489 402390 284694 235555 71050 151221 196107 81543 74709 51795 69769 2509 13088 4001 8052 5-Oxoproline 13C5 19981 6204 9826 17001 419146 169497 336931 161508 18952 22201 52410 28370 10457 29804 14447 14120 75813 120533 52812 95449 14156 16776 37775 25338 7795 8997 8637 11655 472474 723185 631115 475971 566315 206407 447923 543589 491095 595688 360867 424923 82969 83379 84303 103821 S-Adenosyl-L-methionine 13C15 56392 46577 40437 63772 267350 97526 141993 97300 2043743 1296144 569858 686507 447469 160657 450187 75872 132210 353338 414351 36828 988564 1110100 1856033 785774 161001 224651 62340 88473 227752 149565 254776 113139 1463425 159695 1351984 1035462 102407 115472 24302 108385 12520 0 0 17841 Citrulline 13C6 0 4347 0 0 324612 147366 226882 117611 58397 22195 28030 0 3718 0 0 0 103726 146513 182484 102819 5761 42595 33432 60146 0 0 0 5513 333664 524274 451583 322190 202999 105582 171731 244760 251368 331525 232121 299043 37749 31693 40166 49047 Cadaverine 13C5 0 8831 0 0 47114 8181 31535 18911 0 0 4942 0 0 0 0 0 9007 35800 28751 25134 0 0 13320 6758 0 0 0 0 358293 449163 350003 361927 207000 137355 182207 222776 314356 317980 455331 284462 70344 104327 69786 58887 Putrescine 13C4 0 0 0 0 59112 40399 44765 49282 62823 78821 32035 45234 0 0 0 0 34683 41804 8887 51462 47781 114900 89620 48400 0 0 0 0 109742 189663 143748 121345 141214 79325 101673 146307 90475 119079 111760 100608 29417 29658 25536 33915 MS_METABOLITE_DATA_END #METABOLITES METABOLITES_START metabolite_name KEGG ID m/z RT L-alanine C00041 90.0550 0.62 L-arginine C00062 175.1189 0.60 L-asparagine C00152 133.0608 0.62 L-aspartate C00049 134.0449 0.64 L-cysteine C00097 122.0270 0.63 L-glutamate C00025 148.0605 0.63 L-glutamine C00064 147.0764 0.63 L-histidine C00135 156.0770 0.59 L-leucine C00123 132.1019 0.65 L-lysine C00047 147.1129 0.59 L-methionine C00073 150.0580 0.65 L-phenylalanine C00079 166.0863 0.71 L-proline C00148 116.0706 0.64 L-serine C00065 106.0498 0.63 L-threonine C00188 120.0655 0.63 L-tryptophan C00078 205.0972 1.69 L-tyrosine C00082 182.0811 0.65 L-valine C00183 118.0863 0.64 AMP C00020 348.0702 0.66 dAMP C00360 332.0780 0.67 Adenosine C00212 268.1035 0.65 Adenine C00147 136.0619 0.65 GMP C00144 364.0647 0.82 Guanine C00242 152.0562 0.65 CMP C00055 324.0585 0.69 Cytidine C00475 244.0926 0.64 Inosine C00294 269.0878 0.65 Hypoxanthine C00262 137.0459 0.67 Allantoate C00499 177.0609 0.59 5-Hydroxyisourate C11821 185.0310 0.62 2',3'-Cyclic CMP C02354 306.0486 0.66 5-6-Dihydrothymine C00906 129.0659 0.63 Pyridoxal C00250 168.0652 0.64 Nicotinamide C00153 123.0552 0.65 NAD+ C00003 664.1152 0.65 Glutathione C00051 308.0908 0.67 Glutathione disulfide C00127 613.1585 0.82 5-Oxoproline C01879 130.0499 0.63 S-Glutathionyl-L-cysteine C05526 427.0905 0.63 Cys-Gly C01419 179.0479 0.64 gamma-Glutamyl-Se-methylselenocysteine C05695 313.0315 0.60 gamma-L-Glutamyl-D-alanine C03738 219.0972 0.64 gamma-Glutamyl-gamma-aminobutyrate C15767 233.1130 0.65 gamma-L-Glutamylputrescine C15699 218.1503 0.79 (5-L-Glutamyl)-L-glutamine C05283 276.1188 0.64 L-Homocysteine C00155 136.0428 0.64 Cystathionine C00542 223.0742 0.65 Dimethylglycine C01026 104.0706 0.62 Phosphoserine C02532 186.0182 0.61 S-Adenosyl-L-methionine C00019 399.1443 0.59 L-Homoserine C00263 120.0655 0.64 Ornithine C01602 133.0972 0.59 L-Citrulline C00327 176.1029 0.63 Argininosuccinate C03406 291.1294 0.63 Cadaverine C01672 103.1230 0.61 Putrescine C00134 89.1074 0.58 Spermidine C00315 146.1652 0.58 Spermine C00750 203.2229 0.58 Carnosine C00386 227.1136 0.59 5-Guanidino-2-oxopentanoate C03771 174.0873 0.66 Phosphocreatine C02305 212.0427 0.73 Creatine C00300 132.0768 0.63 Creatinine C00791 114.0663 0.63 4-Acetamidobutanoate C02946 146.0811 0.63 N-Acetyl-L-citrulline C15532 218.1142 0.92 N-Acetylornithine C00437 175.1077 0.65 Guanidinoacetate C00581 118.0610 0.62 5-Aminopentanoate C00431 118.0863 0.65 Pantetheine C00831 279.1336 0.65 Pantetheine 4--phosphate C01134 359.1059 1.85 Hypotaurine C00519 110.0270 0.63 L-Methionine S-oxide C02989 166.0521 0.64 5-Hydroxyindoleacetate C05635 192.0622 0.62 3-Methyldioxyindole C05834 146.0601 1.69 Indole C00463 118.0650 0.65 Indole-3-acetaldehyde C00637 160.0751 0.64 Indole-3-acetate C00954 176.0705 1.90 Indolepyruvate C00331 204.0635 0.65 kynurenine C00328 209.0912 0.65 N-formyl kynurenine C02700 237.0902 0.64 Anthranilate C00108 138.0547 0.64 Picolinic acid C10164 124.0394 0.57 g-Oxalo-crotonate C03453 159.0278 0.65 2-Aminomuconate C02220 161.0459 1.90 L-Adrenaline C00788 184.0946 0.62 Dopamine C03758 154.0861 0.64 Serotonin C00780 177.1002 0.63 Acetylcholine C01996 146.1176 0.64 acetyl-carnitine (acyl-C2) C02571 204.1230 0.65 butanoyl-l-carnitine (acyl-C4) C02862 232.1542 0.66 acyl-C4-OH (Hydroxybutyrylcarnitine) HMDB13127 248.1492 0.65 acyl-C5-OH (hydroxyisovalerylcarnitine) ac107 262.1646 0.65 O-dodecanoyl-carnitine (acyl-C12) HMDB02250 344.2792 1.94 O-tetradecanoyl-L-carnitine (acyl-C14) HMDB05066 372.3105 1.99 Tetradecenoyl Carnitine (acyl-C14:1) HMDB13329 370.2949 1.97 L-Palmitoylcarnitine (acyl-C16) C02990 400.3416 2.07 Hexadecenoyl-carnitine (acyl-C16:1) HMDB06317 398.3264 2.01 O-octadecenoyl-L-carnitine (acyl-C18:1) HMDB06351 426.3574 2.09 acyl-C18:2 (Linoleoyl-CoA) HMDB01064 424.3416 2.03 Alanine 13C3 C00041 93.065 0.625515 Arginine 13C6 C00062 181.1391 0.6000713 Asparagine 13C5 C00152 138.0582 0.6336378 Cysteine 13C3 C00097 125.038 0.5613309 Glutamate 13C5 C00025 153.0772 0.643826 Histidine 13C6 C00135 162.0965 0.6293598 Leucine/isoleucine 13C6 C00123 138.1221 0.6467489 Lysine 13C6 C00047 153.1329 0.5781466 Methionine 13C5 C00073 155.0745 0.6357743 Proline 13C5 C00148 121.0871 0.649991 Serine 13C3 C00065 109.0599 0.6274108 Tryptophan 13C11 C00078 216.1366 1.696053 Hypoxanthine 13C5 C00262 142.0622 0.6580078 Xanthine 13C5 C00385 158.0572 0.6606593 5-Oxoproline 13C5 C01879 135.0668 0.638048 S-Adenosyl-L-methionine 13C15 C00019 414.2011 0.6258777 Citrulline 13C6 C00327 182.1234 0.6376935 Cadaverine 13C5 C01672 108.1397 0.577841 Putrescine 13C4 C00134 93.12082 0.5828735 METABOLITES_END #END