#METABOLOMICS WORKBENCH mdm_20250120_221042 DATATRACK_ID:5551 STUDY_ID:ST003683 ANALYSIS_ID:AN006045 PROJECT_ID:PR002283
VERSION             	1
CREATED_ON             	January 24, 2025, 6:38 am
#PROJECT
PR:PROJECT_TITLE                 	Ketogenic diet suppresses colorectal cancer through the gut microbiome long
PR:PROJECT_TITLE                 	chain fatty acid stearate
PR:PROJECT_SUMMARY               	Our manuscript entitled “Ketogenic diet suppresses colorectal cancer through
PR:PROJECT_SUMMARY               	the gut microbiome long chain fatty acid stearate” describes a reduced colonic
PR:PROJECT_SUMMARY               	tumor burden upon ketogenic diet (KD) consumption in a CRC mouse model with a
PR:PROJECT_SUMMARY               	humanized microbiome. Importantly, we demonstrate a causal relationship through
PR:PROJECT_SUMMARY               	microbiome transplantation into germ-free mice, whereby alterations in the gut
PR:PROJECT_SUMMARY               	microbiota were maintained in the absence of continued selective pressure from
PR:PROJECT_SUMMARY               	the KD. Specifically, we identify a shift toward bacterial species that produce
PR:PROJECT_SUMMARY               	stearic acid in ketogenic conditions, whereas consumers were depleted, resulting
PR:PROJECT_SUMMARY               	in elevated levels of free stearate in the gut lumen. This microbial product
PR:PROJECT_SUMMARY               	demonstrates tumor-suppressing properties by inducing apoptosis in cancer cells
PR:PROJECT_SUMMARY               	and decreasing colonic Th17 immune cell populations. As part of this study, we
PR:PROJECT_SUMMARY               	used different metabolomics workflows to study metabolites in mouse fecal and
PR:PROJECT_SUMMARY               	plasma samples as well as the used rodent diet and human feces samples.
PR:INSTITUTE                     	University of Luxembourg
PR:LAST_NAME                     	Letellier
PR:FIRST_NAME                    	Elisabeth
PR:ADDRESS                       	6, avenue du Swing, Belval, Esch, 4367, Luxembourg
PR:EMAIL                         	madita.brauer@uni.lu
PR:PHONE                         	(+352) 46 66 44 6954
#STUDY
ST:STUDY_TITLE                   	Ketogenic diet suppresses colorectal cancer through the gut microbiome long
ST:STUDY_TITLE                   	chain fatty acid stearate - untargeted LCMS data from mouse plasma samples
ST:STUDY_SUMMARY                 	Mouse plasma samples from the SD/KD-fed SPF mice were analyzed by a LC-MS.
ST:STUDY_SUMMARY                 	Metabolites were extracted from human plasma samples as described in the method
ST:STUDY_SUMMARY                 	file. Resulting samples were used for LC-MS untargeted metabolomics screen using
ST:STUDY_SUMMARY                 	a Vanquish UHPLC (ThermoFisher Scientific), coupled to a Q Exactive HF mass
ST:STUDY_SUMMARY                 	spectrometer (ThermoFisher Scientific).
ST:INSTITUTE                     	University of Luxembourg
ST:LAST_NAME                     	Letellier
ST:FIRST_NAME                    	Elisabeth
ST:ADDRESS                       	6, avenue du Swing, Belval, Esch, 4367, Luxembourg
ST:EMAIL                         	madita.brauer@uni.lu
ST:PHONE                         	(+352) 46 66 44 6954
#SUBJECT
SU:SUBJECT_TYPE                  	Mammal
SU:SUBJECT_SPECIES               	Mus musculus
SU:TAXONOMY_ID                   	10090
#SUBJECT_SAMPLE_FACTORS:         	SUBJECT(optional)[tab]SAMPLE[tab]FACTORS(NAME:VALUE pairs separated by |)[tab]Raw file names and additional sample data
SUBJECT_SAMPLE_FACTORS           	KD_11	KD_11	Sample source:Blood plasma | Condition:KD	RAW_FILE_NAME(raw_file_name)=MT001_LC-EMunt_Plasma11_HighFat_LowCarb_1.mzML
SUBJECT_SAMPLE_FACTORS           	KD_12	KD_12	Sample source:Blood plasma | Condition:KD	RAW_FILE_NAME(raw_file_name)=MT001_LC-EMunt_Plasma12_HighFat_LowCarb_1.mzML
SUBJECT_SAMPLE_FACTORS           	KD_13	KD_13	Sample source:Blood plasma | Condition:KD	RAW_FILE_NAME(raw_file_name)=MT001_LC-EMunt_Plasma13_HighFat_LowCarb_1.mzML
SUBJECT_SAMPLE_FACTORS           	SD_16	SD_16	Sample source:Blood plasma | Condition:SD	RAW_FILE_NAME(raw_file_name)=MT001_LC-EMunt_Plasma16_Std-Diet_1.mzML
SUBJECT_SAMPLE_FACTORS           	SD_17	SD_17	Sample source:Blood plasma | Condition:SD	RAW_FILE_NAME(raw_file_name)=MT001_LC-EMunt_Plasma17_Std-Diet_1.mzML
SUBJECT_SAMPLE_FACTORS           	SD_18	SD_18	Sample source:Blood plasma | Condition:SD	RAW_FILE_NAME(raw_file_name)=MT001_LC-EMunt_Plasma18_Std-Diet_1.mzML
SUBJECT_SAMPLE_FACTORS           	SD_19	SD_19	Sample source:Blood plasma | Condition:SD	RAW_FILE_NAME(raw_file_name)=MT001_LC-EMunt_Plasma19_Std-Diet_1.mzML
SUBJECT_SAMPLE_FACTORS           	SD_20	SD_20	Sample source:Blood plasma | Condition:SD	RAW_FILE_NAME(raw_file_name)=MT001_LC-EMunt_Plasma20_Std-Diet_1.mzML
SUBJECT_SAMPLE_FACTORS           	KD_21	KD_21	Sample source:Blood plasma | Condition:KD	RAW_FILE_NAME(raw_file_name)=MT001_LC-EMunt_Plasma21_HighFat_LowCarb_1.mzML
SUBJECT_SAMPLE_FACTORS           	KD_22	KD_22	Sample source:Blood plasma | Condition:KD	RAW_FILE_NAME(raw_file_name)=MT001_LC-EMunt_Plasma22_HighFat_LowCarb_1.mzML
SUBJECT_SAMPLE_FACTORS           	KD_23	KD_23	Sample source:Blood plasma | Condition:KD	RAW_FILE_NAME(raw_file_name)=MT001_LC-EMunt_Plasma23_HighFat_LowCarb_1.mzML
SUBJECT_SAMPLE_FACTORS           	KD_24	KD_24	Sample source:Blood plasma | Condition:KD	RAW_FILE_NAME(raw_file_name)=MT001_LC-EMunt_Plasma24_HighFat_LowCarb_1.mzML
SUBJECT_SAMPLE_FACTORS           	SD_1	SD_1	Sample source:Blood plasma | Condition:SD	RAW_FILE_NAME(raw_file_name)=MT001_LC-EMunt_Plasma01_Std-Diet_1.mzML
SUBJECT_SAMPLE_FACTORS           	SD_2	SD_2	Sample source:Blood plasma | Condition:SD	RAW_FILE_NAME(raw_file_name)=MT001_LC-EMunt_Plasma02_Std-Diet_1.mzML
SUBJECT_SAMPLE_FACTORS           	SD_3	SD_3	Sample source:Blood plasma | Condition:SD	RAW_FILE_NAME(raw_file_name)=MT001_LC-EMunt_Plasma03_Std-Diet_1.mzML
SUBJECT_SAMPLE_FACTORS           	SD_4	SD_4	Sample source:Blood plasma | Condition:SD	RAW_FILE_NAME(raw_file_name)=MT001_LC-EMunt_Plasma04_Std-Diet_1.mzML
#COLLECTION
CO:COLLECTION_SUMMARY            	Plasma samples from SPF mice fed a ketogenic or standard diet were taken and
CO:COLLECTION_SUMMARY            	stored in plasma collection tube at -80℃ until use for metabolite
CO:COLLECTION_SUMMARY            	extraction.
CO:SAMPLE_TYPE                   	Blood (plasma)
CO:STORAGE_CONDITIONS            	-80℃
#TREATMENT
TR:TREATMENT_SUMMARY             	Mice were subjected to AOM/DSS treatment and received either a ketogenic diet
TR:TREATMENT_SUMMARY             	(KD) or a standard diet (SD) as described in Tsenkova et al. (2025). Briefly,
TR:TREATMENT_SUMMARY             	mice were administered a single intraperitoneal injection of sterile-filtered
TR:TREATMENT_SUMMARY             	AOM (10 mg/kg mouse body weight). SPF mice were administered a sterile-filtered
TR:TREATMENT_SUMMARY             	antibiotic mix (vancomycin hydrochloride (Sigma-Aldrich, 500 mg/L), ampicillin
TR:TREATMENT_SUMMARY             	(Sigma-Aldrich, 1 g/L), neomycin (Sigma-Aldrich, 1 g/L) and metronidazole
TR:TREATMENT_SUMMARY             	(Sigma-Aldrich, 1 g/L)) via their drinking water for one week prior to AOM
TR:TREATMENT_SUMMARY             	injection. Microbiomes were replaced with donor microbiomes via oral gavage.
TR:TREATMENT_SUMMARY             	Mice were administered a total of three cycles of DSS, with a two-week-long
TR:TREATMENT_SUMMARY             	recovery period between each cycle. DSS was refreshed mid-cycle. Half of the
TR:TREATMENT_SUMMARY             	mice were randomly allocated to each dietary condition at the end of the last
TR:TREATMENT_SUMMARY             	cycle of DSS.
#SAMPLEPREP
SP:SAMPLEPREP_SUMMARY            	Plasma samples were thawed on ice and centrifuged at 15000g for three minutes at
SP:SAMPLEPREP_SUMMARY            	4°C. An ISM was prepared (1 mg/mL of 6-chloropurine riboside,
SP:SAMPLEPREP_SUMMARY            	2-chloroquinoline-3-carboxylic acid, 4-chloro-DL-phenylalanine,
SP:SAMPLEPREP_SUMMARY            	Nε-trifluoroacetyl-L-lysine, sucralose, caffeine-trimethyl in MilliQ® water).
SP:SAMPLEPREP_SUMMARY            	20 μL of the ISM was added to 20 μL of plasma at 4°C and the samples were
SP:SAMPLEPREP_SUMMARY            	vortexed thoroughly. Proteins were precipitated by the addition of 151 μL of
SP:SAMPLEPREP_SUMMARY            	methanol to 38 μL of sample at 4°C and vortexed thoroughly. Samples were
SP:SAMPLEPREP_SUMMARY            	incubated for 15 minutes at -20°C, then centrifuged for five minutes at 4°C in
SP:SAMPLEPREP_SUMMARY            	a ThermoMixer. Phospholipids were removed by diluting samples in 300μL of
SP:SAMPLEPREP_SUMMARY            	methanol, then transferring them to a Phree phospholipid removal plate and
SP:SAMPLEPREP_SUMMARY            	vacuum for 2-7 inches Hg was applied until the filtrate collected in the deep
SP:SAMPLEPREP_SUMMARY            	well plate. 350 μL were transferred into 1.5mL Eppendorf tubes and the solvents
SP:SAMPLEPREP_SUMMARY            	were evaporated in a SpeedVac® at 4°C overnight. Then, the temperature of the
SP:SAMPLEPREP_SUMMARY            	SpeedVac® was increased to room temperature for 25 minutes to avoid water
SP:SAMPLEPREP_SUMMARY            	condensation on the surface of the tube. Samples were stored at -80°C until
SP:SAMPLEPREP_SUMMARY            	LC-MS analysis was performed.
#CHROMATOGRAPHY
CH:CHROMATOGRAPHY_TYPE           	HILIC
CH:INSTRUMENT_NAME               	Thermo Vanquish
CH:COLUMN_NAME                   	SeQuant ZIC- pHILIC (150 x 2.1 mm, 5 μm)
CH:SOLVENT_A                     	100% Water; 0.01% formic acid
CH:SOLVENT_B                     	90% Acetonitrile/10% Water; 0.1% formic acid; 20 mM ammonium acetate
CH:FLOW_GRADIENT                 	90 % B for 1.5 min, followed by a decrease to 20% B within 15 min, then 20% B
CH:FLOW_GRADIENT                 	for 2 min, increase to 90% B within 2 min, then 90% B for 13 min
CH:FLOW_RATE                     	200 µL/mL
CH:COLUMN_TEMPERATURE            	50°C
#ANALYSIS
AN:ANALYSIS_TYPE                 	MS
#MS
MS:INSTRUMENT_NAME               	Thermo Q Exactive HF-X Orbitrap
MS:INSTRUMENT_TYPE               	Orbitrap
MS:MS_TYPE                       	ESI
MS:ION_MODE                      	UNSPECIFIED
MS:MS_COMMENTS                   	Resolution - 120,000, m/z range - 60-900, AGC target - 1e6, maximum injection
MS:MS_COMMENTS                   	time – 70 ms. ddMS2 was applied using the following settings: Resolution -
MS:MS_COMMENTS                   	30,000, AGC target - 5e5, maximum injection time – 70, topN - 5, Normalized
MS:MS_COMMENTS                   	collision energy – 20. Samples were acquired in positive and negative
MS:MS_COMMENTS                   	ionization mode simultaneously (polarity switching).
#MS_METABOLITE_DATA
MS_METABOLITE_DATA:UNITS	area
MS_METABOLITE_DATA_START
Samples	SD_1	SD_2	SD_3	SD_4	KD_11	KD_12	KD_13	SD_16	SD_17	SD_18	SD_19	SD_20	KD_21	KD_22	KD_23	KD_24
Factors	Sample source:Blood plasma | Condition:SD	Sample source:Blood plasma | Condition:SD	Sample source:Blood plasma | Condition:SD	Sample source:Blood plasma | Condition:SD	Sample source:Blood plasma | Condition:KD	Sample source:Blood plasma | Condition:KD	Sample source:Blood plasma | Condition:KD	Sample source:Blood plasma | Condition:SD	Sample source:Blood plasma | Condition:SD	Sample source:Blood plasma | Condition:SD	Sample source:Blood plasma | Condition:SD	Sample source:Blood plasma | Condition:SD	Sample source:Blood plasma | Condition:KD	Sample source:Blood plasma | Condition:KD	Sample source:Blood plasma | Condition:KD	Sample source:Blood plasma | Condition:KD
2-Hydroxybutyric acid	119921629	111321348	134888672	107454910	254686564	250894670	214741003	85947376	85432893	70166964	67563578	75488636	119978728	131809150	101128333	100525691
2-Hydroxyglutaric acid	37571324	29280401	34034814	29905584	22750139	30493163	21011478	29233121	17881564	21889609	24920511	29544416	15486496	20491456	16480671	21854114
3-(4-Hydroxyphenyl)propionic acid	5105543	5858624	6517016	6183801	5121079	5397627	6101729	8962026	6351606	7544784	9561951	7918218	5069001	6057993	5712840	5847349
3-Aminobutanoic acid	2879307	2549618	27494012	2811632	4492024	2961203	4035489	14965286	2508216	14982724	2279439	2500574	3706615	4563208	2382259	4885473
3-Hydroxyanthranilic acid	18040724	32829130	20748018	5454904	20798677	5665581	3946265	4923148	16148885	25388240	8881604	4002156	990454	8686822	2988149	3123540
3-Methoxytyramine	2567142	3281862	3786154	3431358	3354682	2923200	2792634	2683063	2969596	2810952	3042214	2603146	1450885	2737149	3096388	2208888
Acetylcarnitine	1910940529	1065767867	766186065	710874611	956191108	872529171	764760310	1398440639	985311045	803882460	577413472	1103090469	991844751	492596207	576682105	333974677
Aconitic Acid	17750595	17161408	22937786	21796435	21629400	20073057	17852815	26925657	17167981	20889811	21782163	23071986	18535578	22012579	15725101	17471914
Adenine	4935037	4591962	4348285	4034026	3025788	4718219	3203535	2573180	2396163	2489068	2755255	2722232	3177931	2134821	2400669	2529826
Alanine	1419667176	952582650	1394804333	1188242842	1359970762	1252692254	1361860724	1354189654	1315761954	1196543740	1253088036	1635659221	994791928	887845576	814896476	1256023101
Phenylalanine	403778656	304866559	315030671	184031101	358481399	413350156	409190264	381077857	379058992	460018490	391344504	431570464	427136661	326887154	388702569	387326696
alpha-Ketoglutaric acid	17060271	19093507	14236751	17757111	13918978	14979115	14722149	12829601	22821931	16236091	18901621	15393880	17842288	18548606	25340819	22212592
Anthranilic acid	35712	48079	29537	31831	38861	26373	37378	93037	99829	66797	76035	69124	103912	154154	230414	53931
Arginine	203320420	190639592	217045241	206164149	163826630	84215888	186873823	219890689	77556243	160155135	206861444	204904032	5269845	58110655	233987918	180619575
Asparagine	31085742	25481927	37302155	22483503	31125125	36407432	36061257	35396736	28099470	28730230	24327956	36815979	41694870	26621829	32601874	53953492
Aspartic acid	67690817	54794994	111487084	80789386	51620017	40634643	46654726	56756396	49761737	87511462	42089267	44388113	85254873	45868962	37303488	48467050
N-Acetylaspartic acid	22421440	17997558	22947320	25986682	27595362	27162238	26176813	19381477	15073579	19070110	20991900	19891127	22089767	19508026	14098691	15757579
Betaine	291355054	315678504	380818019	240518134	414004029	352471538	399269728	355422000	367633243	364748640	375817407	373607473	450659677	315650823	432070499	557137820
Carnitine	3547143269	2734071036	2399506505	2503392934	2063072971	1976487405	2080243345	2861202208	2587701051	2630989244	2150926353	2553163523	1399046275	1329872280	1090793640	1268245363
Myristoylcarnitine	112358	394891	142189	429505	1346150	1563264	585042	344788	319581	787796	298775	16170	622423	786667	880847	517575
Choline	3054459008	2669249124	2921527908	3048474649	2855067530	2963331399	2529028994	2478828443	2263097120	2440442073	2264208820	2518280064	2996069544	2502452763	2519047347	2353054423
Citric acid	186110535	366066495	415042156	531738892	603989409	514725196	305811004	549837899	227986476	351108221	446554001	480889429	336269875	598355565	125475270	241429150
Isocitric acid	187891970	367539769	416783794	533707145	606035715	517040543	308098780	552004870	231011349	353100156	448802971	482873973	339276858	601129517	128707518	243994360
Citrulline	227939116	207114179	294671286	311453938	308729161	300396931	321854644	265413113	240772745	332460433	273079671	267913316	327378510	324320666	256920819	302695237
Creatine	4493449082	3713309996	4103891286	3666422690	4103100499	3686521961	4072421816	3331405919	4198943317	3492024075	3095245836	3661229426	4132628810	3244913421	3958240238	2643905141
Creatinine	1051371986	866930531	888064151	834146293	790920867	885009231	983534546	898013444	1020346291	1034296945	1010257355	1078458016	704694524	796602856	622708917	663475045
Cystine	4205822	1731004	3542856	3609661	7102663	7011726	5597232	3665506	5136274	3989561	3658237	4053998	4916353	6998646	8644764	7578598
Cytosine	10982981	13727861	7621735	7972894	17742067	18601025	25112564	13358023	14251445	17411699	15250607	12159458	15934897	15590291	14514402	15074631
Dopamine	4217340	4965112	4438109	3990309	3408753	2764027	1560234	4385415	5076525	3787544	5445762	5351702	2089259	3118682	2546851	2395250
D-PANTOTHENIC ACID	4043156	2061597	2599860	1767912	2076757	4163520	1888256	9027246	5171688	5398498	5260220	8495116	5395750	5385305	3016561	3505061
Fructose	186021072	237987563	312018501	313290265	447308260	522594056	452044078	212090475	451012396	438513087	396321701	330384560	659541891	530953912	545071632	493271769
Fructose 6-phosphate	1253890	1095282	790425	1645696	1991704	2547933	880061	1085816	1644144	927814	639628	541340	9580074	4060788	1149738	803010
Fumaric acid	36405208	31095498	55522141	36815920	43943973	39324240	35408067	60749854	15594802	33984499	27987448	48340339	25182362	24958236	10977929	16597973
Galactose	185007959	224314693	299168183	300714722	440397024	511627559	442216134	201034448	440014038	427055342	386814059	329104257	650020749	521170550	538961838	483070847
gamma-Aminobutyric acid	298426313	229606553	535556284	249385022	229338171	211634604	253917366	285747870	309234824	282744818	211263116	255120100	216830022	292894707	137722205	182955276
Glucose	185007959	224314693	299168183	300714722	440397024	511627559	442216134	201034448	440014038	427055342	386814059	329104257	650020749	521170550	538961838	483070847
Glucose 1-phosphate	2214553	13733765	2513036	1505318	15223611	5891972	3046790	829395	1373531	482142	426165	263027	9514019	4873433	921703	643470
Glucose-6-phosphate	2214553	13733765	2513036	1505318	15223611	5891972	3046790	829395	1373531	482142	426165	263027	9514019	4873433	921703	643470
Glutamic acid	69505078	78983292	85521021	83817669	78996128	73064620	47018921	49913767	62316278	57836254	38330733	43348752	102788467	70454438	46377511	43747359
Glutamine	2258445899	1736588041	2076056410	1902113363	1775599090	1977784563	2220298275	2171395367	2302557813	2193535143	2056238407	2248868288	2078746926	1783506879	1949081183	1851928935
Glutathione, oxidized (GSSG)	321193	366601	402388	790873	663744	895870	98518	481320	224877	99691	227015	429450	2116040	1986540	7845	199768
Glycine	40804389	29992991	55127758	46354696	29256514	25928362	27686813	28095542	27340121	26846001	29561346	38306568	28768733	23387476	20705728	22294490
Isobutyrylglycine	20079295	11154138	41578080	13154800	16061403	10390127	12817687	10088594	4254672	6025802	5611932	9290248	5523382	4756820	5871906	4594263
HIAA	649388	959110	903145	1181806	537213	470992	574109	639685	1164604	1078276	1231512	735901	705276	396562	617715	599438
Histidine	623118845	434267224	528310264	316305918	770014620	706721231	806779981	762944308	934998026	914493299	916748199	975985968	1059284367	663416897	853334435	986215265
S-Adenosyl-L-homocysteine, SAH	181000	344627	472030	286266	140885	306252	46186	245367	1828797	586589	114344	325092	703142	944730	226173	144649
Hydroxyproline	31612382	35595724	58057145	57132769	27289792	29012089	31493378	25127918	27583293	27543236	53992912	45946540	10444754	14033909	8724673	10516639
3-Methylindole	2356230	3211467	3408370	1332386	1671422	2638302	4272079	10027890	9491762	10938190	3943057	12099552	2851530	2110303	3020866	8924938
Indole	1442599	3845804	3522636	3050389	916197	1730946	1215707	5096624	5534166	5960468	4641952	5242939	2130737	1517261	2087011	3187953
Indole-3-acetaldehyde	18158340	18621376	4439695	2540378	7893210	61779511	25955592	15699390	35582488	15802247	19530886	23275492	46499822	34924226	34717992	11619858
Indole-3-acrylic acid	66780071	88003871	77964075	73302735	51215830	70284257	63157420	99757820	125681357	122255339	101895077	113396705	82210020	79347386	83521494	96401623
Indole-3-carboxaldehyde	9944302	15651874	9034672	11548111	8638835	15461598	10244533	13608255	21110424	29555174	19398526	14214279	30679974	22337304	35917274	20224640
Indole-3-carboxylic acid	113318	245424	81198	110806	166016	92015	126380	22196	71134	94068	51839	67064	96848	188053	107888	33836
Indole-3-glyoxylic acid	57380694	71707213	82480047	39353686	34442944	72883780	43306094	72212952	71015826	72948595	82298878	74596305	70542487	68790778	77036824	75307920
Indole-3-lactic acid	1722745	2373666	2382944	2399546	2613393	3086260	2718705	4046711	4270433	5626849	4126412	3747074	4493392	3707506	4165914	3732259
Indole-3-propionic acid	2128577	12849062	45458340	29899432	2175214	2562247	2524823	64170852	15173497	23270657	46828998	26355220	2443016	2960784	838360	3448376
Isoleucine	833082975	775790441	919657450	583444353	1062676444	957285018	983690913	1136085883	1206704749	1079954155	1004851422	1122372877	1630945814	948458906	1356810653	1264545324
Kynurenic acid	33001216	39710273	44925713	44311966	39163788	41071134	25828520	40090539	39616404	40734758	44242692	41248939	37997306	38658682	42179455	41938596
Kynurenine	3162685	4515384	4267947	6072981	2405631	2711687	3087053	3218524	5341243	4598320	6128335	3571044	3478380	2363565	3149951	2873340
Lactic Acid	4481970192	2379354006	2321944458	1786355998	2345034498	3044060168	2027660384	3691689830	2533765873	2018259119	1945087810	3007151868	1782622809	1738174090	1016083940	1364308202
Leucine	833082975	775790441	919657450	583444353	1062676444	957285018	983690913	1136085883	1206704749	1079954155	1004851422	1122372877	1630945814	948458906	1356810653	1264545324
Lysine	188334951	116714049	278537639	116691130	233319092	229166769	280293967	221221286	242234454	201065760	188696339	262940217	186877783	161592627	208633660	247866808
Malic Acid	384564271	323363410	604415858	394700388	471731209	414718884	369492464	657640688	143192207	364449803	281962928	509130079	250810595	248806920	83983503	149899262
Mannose	185007959	224314693	299168183	300714722	440397024	511627559	442216134	201034448	440014038	427055342	386814059	329104257	650020749	521170550	538961838	483070847
Mannose 1-phosphate	2214553	13733765	2513036	1505318	15223611	5891972	3046790	829395	1373531	482142	426165	263027	9514019	4873433	921703	643470
Methylmalonic acid	80936153	110051679	99840019	114653629	122064153	104266003	73133771	100835705	51282866	81291331	86882764	108924886	43545022	96117591	39324907	100851590
Niacin	309994	497054	358428	384086	433931	582005	402640	252132	287521	379865	401350	392464	413695	580066	585327	693935
Nicotinamide	38552477	24763034	31808663	38747954	42011215	63234116	36698674	39691958	57411780	49093046	57035803	44533736	138785101	79435745	87723004	99673428
Dihydronicotinamide riboside	48705	37648	117000	99927	154886	316360	53495	222168	95199	337504	172474	187250	237480	128465	201579	225518
Nicotinamide riboside, NR	4851071	10897134	17695516	12991331	7523240	7976142	7262790	13626772	16766455	18705973	18485653	18209242	19615798	12718322	16209242	22274791
Ornithine	174703545	151648500	280051972	189209522	172625557	253864563	201583968	209039066	291142841	264119353	206670764	258620341	326214260	302539115	195758634	220878491
Proline	802207029	557395177	962394407	700372567	836137906	750934615	903119304	658589703	644754944	611284009	642592196	995278189	640697206	482620656	611678960	1094901218
Putrescine	1804865249	1795613319	1608429354	1534475339	1343303196	1355822835	1310225047	1293809723	1351257283	1337079186	1356438057	1335270678	1307292822	1302630513	1382594419	1380811724
Quinaldic acid	52669	87720	63853	69282	134212	56682	53568	83282	86218	172110	71112	111348	36140	91634	75852	78136
Quinolinic acid	504746	590240	228648	102537	135319	341939	370428	94257	433123	333069	108957	80182	556989	433810	459964	566577
Sarcosine	2729254	3418264	12542990	5009893	5704281	5002644	5837031	6616503	4734428	5861663	4488693	5891255	4699819	4632698	2862543	4847395
Serine	36706253	34297908	37911679	24044610	46715368	47306160	51423344	40660782	47254632	45404308	42933946	53931227	54577027	45330338	42588061	60258265
Serotonin	4650020	5072495	932305	202753	1625261	16427276	6701203	3973388	10153656	3717038	5712690	6212787	13510955	8458052	8790691	2578569
Succinic acid	187897426	289197402	229058455	277032354	307079894	250558064	167842349	239850373	102517767	189905920	205508410	256627480	83911468	228972855	74005980	237846901
Taurine	483734314	349776284	297521071	225955164	492467239	623159035	463069141	358214468	299403800	398287197	247869188	303816031	702289283	400203697	371939311	340233044
Threonine	251179446	195866141	328582235	279943406	342976848	315975415	382437838	253294171	269559926	247237115	271696237	336393081	285442384	249514863	232232388	414970898
Thymine	37328318	21286613	44874114	38935700	36836424	32505205	44786136	49161249	33253980	57403364	38079374	58722126	28698996	36643850	33045126	50761222
Tryptophan	77508487	103423214	90567142	85916023	59004461	80134722	72600497	114597221	144075518	138528786	114144357	127519123	93253005	89769217	94846996	106086172
Tyrosine	153898041	206992437	289829219	227503215	315649415	377449136	350638802	181695321	307360510	330748984	278904268	294585868	435758344	358533604	398737787	332844520
Uracil	10767030	3383591	9954605	5327750	10075709	6941816	9310950	6763179	5724290	5324425	2640747	4348904	13670141	5326290	4813832	3807267
Uridine	9932380	8679118	10361412	11341718	10040312	9176730	10266194	8964108	9673066	10049981	7905232	9458042	10902311	9241744	8409868	6890268
Valine	6079955596	4007097731	8191115713	6119569891	3991574241	3624510920	4288388680	5793752750	5579522514	4661119896	4108710213	5264897826	4407112495	3910261079	3247378162	3313926106
MS_METABOLITE_DATA_END
#METABOLITES
METABOLITES_START
metabolite_name
2-Hydroxybutyric acid
2-Hydroxyglutaric acid
3-(4-Hydroxyphenyl)propionic acid
3-Aminobutanoic acid
3-Hydroxyanthranilic acid
3-Methoxytyramine
Acetylcarnitine
Aconitic Acid
Adenine
Alanine
Phenylalanine
alpha-Ketoglutaric acid
Anthranilic acid
Arginine
Asparagine
Aspartic acid
N-Acetylaspartic acid
Betaine
Carnitine
Myristoylcarnitine
Choline
Citric acid
Isocitric acid
Citrulline
Creatine
Creatinine
Cystine
Cytosine
Dopamine
D-PANTOTHENIC ACID
Fructose
Fructose 6-phosphate
Fumaric acid
Galactose
gamma-Aminobutyric acid
Glucose
Glucose 1-phosphate
Glucose-6-phosphate
Glutamic acid
Glutamine
Glutathione, oxidized (GSSG)
Glycine
Isobutyrylglycine
HIAA
Histidine
S-Adenosyl-L-homocysteine, SAH
Hydroxyproline
3-Methylindole
Indole
Indole-3-acetaldehyde
Indole-3-acrylic acid
Indole-3-carboxaldehyde
Indole-3-carboxylic acid
Indole-3-glyoxylic acid
Indole-3-lactic acid
Indole-3-propionic acid
Isoleucine
Kynurenic acid
Kynurenine
Lactic Acid
Leucine
Lysine
Malic Acid
Mannose
Mannose 1-phosphate
Methylmalonic acid
Niacin
Nicotinamide
Dihydronicotinamide riboside
Nicotinamide riboside, NR
Ornithine
Proline
Putrescine
Quinaldic acid
Quinolinic acid
Sarcosine
Serine
Serotonin
Succinic acid
Taurine
Threonine
Thymine
Tryptophan
Tyrosine
Uracil
Uridine
Valine
METABOLITES_END
#END