#METABOLOMICS WORKBENCH shancock_20250221_204759 DATATRACK_ID:5666 STUDY_ID:ST003764 ANALYSIS_ID:AN006179 PROJECT_ID:PR002347 VERSION 1 CREATED_ON February 21, 2025, 9:24 pm #PROJECT PR:PROJECT_TITLE Profiling of fatty acid double bond positing ing gemcitabine-resistant PR:PROJECT_TITLE pancreatic ductal adenocarcinoma cells PR:PROJECT_SUMMARY Pancreatic ductal adenocarcinoma (PDAC) is a highly aggressive disease with few PR:PROJECT_SUMMARY treatment options and poor survivability. In this work we sought to characterise PR:PROJECT_SUMMARY metabolic adaptations to gemcitabine (GEMC)-based chemotherapy exposure to PR:PROJECT_SUMMARY discover new therapeutic targets for improving treatment efficacy. We show that PR:PROJECT_SUMMARY GEMC resistance (GEMR) upregulates de novo lipogenesis in Panc1 and MiaPaCa2 PR:PROJECT_SUMMARY cells. PR:INSTITUTE Victor Chang Cardiac Research Institute PR:LAST_NAME Hancock PR:FIRST_NAME Sarah PR:ADDRESS Level 7 Lowy Packer Building, 405 Liverpool Street, Darlinghurst, NSW 2010, PR:ADDRESS Australia PR:EMAIL s.hancock@victorchang.edu.au PR:PHONE +61414537526 #STUDY ST:STUDY_TITLE Profiling of fatty acid double bond positing ing gemcitabine-resistant ST:STUDY_TITLE pancreatic ductal adenocarcinoma cells ST:STUDY_SUMMARY In this work we developed gemcitabine (GEMC) resistant (GEMR) PDAC cell lines ST:STUDY_SUMMARY from Panc1 and MiaPaCa2 cells. This dataset contains ozone-induced fatty acid ST:STUDY_SUMMARY discovery (OzFAD) data, which shows differences in carbon-carbon double bond ST:STUDY_SUMMARY position between Panc1 and MiaPaCa2 cells. Panc1 cells contain significant ST:STUDY_SUMMARY amounts of 16:1n-10, which is an unusual isomeric fatty acid produced by Δ6 ST:STUDY_SUMMARY desaturation of palmitic acid by fatty acid desaturase 2 (FADS2). Panc1 GEMR ST:STUDY_SUMMARY cells show increased stearoyl-CoA desaturase 1 (SCD1)-derived 16:1n-7 alongside ST:STUDY_SUMMARY decreased 16:1n-10. ST:INSTITUTE Victor Chang Cardiac Research Institute ST:LAST_NAME Hancock ST:FIRST_NAME Sarah ST:ADDRESS Level 7 Lowy Packer Building, 405 Liverpool Street, Darlinghurst, NSW 2010, ST:ADDRESS Australia ST:EMAIL s.hancock@victorchang.edu.au ST:PHONE +61414537526 #SUBJECT SU:SUBJECT_TYPE Cultured cells SU:SUBJECT_SPECIES Homo sapiens SU:TAXONOMY_ID 9606 #SUBJECT_SAMPLE_FACTORS: SUBJECT(optional)[tab]SAMPLE[tab]FACTORS(NAME:VALUE pairs separated by |)[tab]Raw file names and additional sample data SUBJECT_SAMPLE_FACTORS - M64_1 Sample source:MiaPaCa2 cells | group:GEMR RAW_FILE_NAME(Raw file name)=20240221_MS_5uL_M64_1.mzML SUBJECT_SAMPLE_FACTORS - M64_2 Sample source:MiaPaCa2 cells | group:GEMR RAW_FILE_NAME(Raw file name)=20240221_MS_5uL_M64_2.mzML SUBJECT_SAMPLE_FACTORS - M64_3 Sample source:MiaPaCa2 cells | group:GEMR RAW_FILE_NAME(Raw file name)=20240221_MS_5uL_M64_3.mzML SUBJECT_SAMPLE_FACTORS - MIA_1 Sample source:MiaPaCa2 cells | group:CON RAW_FILE_NAME(Raw file name)=20240221_MS_5uL_MIA_1.mzML SUBJECT_SAMPLE_FACTORS - MIA_2 Sample source:MiaPaCa2 cells | group:CON RAW_FILE_NAME(Raw file name)=20240221_MS_5uL_MIA_2.mzML SUBJECT_SAMPLE_FACTORS - MIA_3 Sample source:MiaPaCa2 cells | group:CON RAW_FILE_NAME(Raw file name)=20240221_MS_5uL_MIA_3.mzML SUBJECT_SAMPLE_FACTORS - P150_1 Sample source:Panc1 cells | group:GEMR RAW_FILE_NAME(Raw file name)=20240221_MS_5uL_P150_1.mzML SUBJECT_SAMPLE_FACTORS - P150_2 Sample source:Panc1 cells | group:GEMR RAW_FILE_NAME(Raw file name)=20240221_MS_5uL_P150_2.mzML SUBJECT_SAMPLE_FACTORS - P150_3 Sample source:Panc1 cells | group:GEMR RAW_FILE_NAME(Raw file name)=20240221_MS_5uL_P150_3.mzML SUBJECT_SAMPLE_FACTORS - PANC_1 Sample source:Panc1 cells | group:CON RAW_FILE_NAME(Raw file name)=20240221_MS_5uL_PANC_1.mzML SUBJECT_SAMPLE_FACTORS - PANC_2 Sample source:Panc1 cells | group:CON RAW_FILE_NAME(Raw file name)=20240221_MS_5uL_PANC_2.mzML SUBJECT_SAMPLE_FACTORS - PANC_3 Sample source:Panc1 cells | group:CON RAW_FILE_NAME(Raw file name)=20240221_MS_5uL_PANC_3.mzML #COLLECTION CO:COLLECTION_SUMMARY Samples were obtained from established immortalised adherent human pancreatic CO:COLLECTION_SUMMARY ductal adenocarcinoma cells (Panc1). Cells were cultured in high glucose DMEM CO:COLLECTION_SUMMARY containing 4.5 g/L glucose and 4 mM glutamine and were supplemented with 10% CO:COLLECTION_SUMMARY FCS. Cells were cultured under 5% CO2 at 37ºC and routinely passaged every 2-3 CO:COLLECTION_SUMMARY days (when ~80-90% confluent). Cell lines were regularly screened for mycoplasma CO:COLLECTION_SUMMARY infection. CO:SAMPLE_TYPE Pancreas #TREATMENT TR:TREATMENT_SUMMARY Panc1 GEMR cells were produced by serial treatment with escalating doses of TR:TREATMENT_SUMMARY gemcitabine for approximately 12 weeks. The final concentration of GEMC for TR:TREATMENT_SUMMARY Panc1 cells was 150 nM, resulting in a half-maximal inhibitory concentration of TR:TREATMENT_SUMMARY 720.5 nM (approximately 2.5 times increase over control (CON) cells, 273.3 nM). TR:TREATMENT_SUMMARY The final concentration of GEMC for MiaPaCa2 cells was 64 nM, resulting in a TR:TREATMENT_SUMMARY half-maximal inhibitory concentration of 92.2 nM (approximately 3x increase over TR:TREATMENT_SUMMARY control (CON) cells, 30.6 nM). #SAMPLEPREP SP:SAMPLEPREP_SUMMARY Lipids were extracted from cells grown to 80-90% confluency in 6 well plates SP:SAMPLEPREP_SUMMARY using a modified methyl tert-butyl ether (MTBE) method. Adherent cells were SP:SAMPLEPREP_SUMMARY washed with ice-cold PBS and scraped into methanol containing 0.01% butylated SP:SAMPLEPREP_SUMMARY hydroxytoluene and 100 pmol each of internal standards (phosphatidylcholine SP:SAMPLEPREP_SUMMARY 19:0/19:0, phosphatidylethanolamine 17:0/17:0, phosphatidylserine 17:0/17:0, SP:SAMPLEPREP_SUMMARY lysophosphatidylcholine 17:0, lysophosphatidylethanolamine 14:0, SP:SAMPLEPREP_SUMMARY phosphatidylglycerol 17:0/17:0, phosphatidic acid 17:0/17:0, ceramide SP:SAMPLEPREP_SUMMARY d18:1/17:0, dihydrosphingomyelin 12:0, cholesteryl ester 22:1, diacylglycerol SP:SAMPLEPREP_SUMMARY 17:0/17:0, D5-triacylglycerol 16:0/16:0/16:0 and cardiolipin SP:SAMPLEPREP_SUMMARY 14:0/14:0/14:0/14:0, and 20 nmol of methyl nonadecanoate). An empty well was SP:SAMPLEPREP_SUMMARY also scraped for use in background subtraction. To this, MTBE was added at a SP:SAMPLEPREP_SUMMARY ratio of 3:1 v/v, and extracts were rotated overnight at 4ºC. Following SP:SAMPLEPREP_SUMMARY overnight rotation, 1 part of ice-cold 150 mM ammonium acetate was added, SP:SAMPLEPREP_SUMMARY samples were vortexed vigorously and centrifuged at 2000 x g for 5 min. The SP:SAMPLEPREP_SUMMARY upper organic phase containing lipids was transferred into a 1.5 mL autosampler SP:SAMPLEPREP_SUMMARY vial and dried under N2 at 37ºC Dried lipids were reconstituted in SP:SAMPLEPREP_SUMMARY chloroform:methanol:water (60:30:4.5 v/v/v), transferred to a sleeved vial, and SP:SAMPLEPREP_SUMMARY stored at -20ºC until analysis. The aqueous phase containing the protein pellet SP:SAMPLEPREP_SUMMARY was dried under N2 and digested with 1M NaOH overnight at 4ºC. Digested protein SP:SAMPLEPREP_SUMMARY was diluted 1:2 v/v with water and protein concentration was determined using a SP:SAMPLEPREP_SUMMARY Pierce BCA assay kit (ThermoFisher Scientific) Following MTBE extraction of SP:SAMPLEPREP_SUMMARY lipids, an aliquot was taken and hydrolysed in 0.6 M KOH in 75% v/v methanol at SP:SAMPLEPREP_SUMMARY 60ºC for 30 min. Samples were then cooled to room temperature and neutralised SP:SAMPLEPREP_SUMMARY with 25% v/v acetic acid. Water and n-hexane (3:2 v/v) were added separately and SP:SAMPLEPREP_SUMMARY samples were vortexed vigorously before being centrifuged at 2000 x g for 5 min. SP:SAMPLEPREP_SUMMARY The upper phase containing hydrolysed fatty acids was removed, and the aqueous SP:SAMPLEPREP_SUMMARY phase was washed with a second volume of n-hexane and combined. The combined SP:SAMPLEPREP_SUMMARY organic phase was then dried under N2 at 37°C. Dried hydrolysed fatty acids SP:SAMPLEPREP_SUMMARY were derivatised with an AMP+ MaxSpec kit (Cayman Chemical, Ann Arbor, MI, USA) SP:SAMPLEPREP_SUMMARY as per manufacturer instructions. Following this, AMP-derivatised fatty acids SP:SAMPLEPREP_SUMMARY were re-extracted using MTBE:water (1:1 v/v), and the upper MTBE phase was dried SP:SAMPLEPREP_SUMMARY under N2 before being resuspended in methanol. Samples were stored at -20°C SP:SAMPLEPREP_SUMMARY until analysis. A 37-component fatty acid methyl ester standard (Merck Life SP:SAMPLEPREP_SUMMARY Sciences) containing an additional 3.184 nmol of methyl nonadecanoate was SP:SAMPLEPREP_SUMMARY concurrently subjected to the same derivatisation procedure and used as an SP:SAMPLEPREP_SUMMARY external quality control for fatty acid identification by LC-MS. #CHROMATOGRAPHY CH:CHROMATOGRAPHY_TYPE Reversed phase CH:INSTRUMENT_NAME Waters Acquity CH:COLUMN_NAME Waters ACQUITY UPLC CSH C18 (100 x 2.1mm,1.7um) CH:SOLVENT_A Water (0.1% formic acid) CH:SOLVENT_B Acetonitrile (0.1% formic acid) CH:FLOW_GRADIENT 0 - 0.5 min at 20% B, then linearly increased to 90% B (16 min), then 100% B CH:FLOW_GRADIENT (17.35 min), held at 100% B until 20.35 min, reduced to 20% B and held until 21 CH:FLOW_GRADIENT min. CH:FLOW_RATE 0.4 mL/min CH:COLUMN_TEMPERATURE 60 #ANALYSIS AN:ANALYSIS_TYPE MS #MS MS:INSTRUMENT_NAME Waters Synapt G2 Si QTOF MS:INSTRUMENT_TYPE QTOF MS:MS_TYPE ESI MS:ION_MODE POSITIVE MS:MS_COMMENTS Chromatographic detection was conducted using an ion-mobility enabled quadrupole MS:MS_COMMENTS time of flight (QTOF) mass spectrometer (SYNAPT G2-Si HDMS, Waters Corporation) MS:MS_COMMENTS previously modified to allow introduction of high concentration ozone into the MS:MS_COMMENTS ion mobility region of the instrument 87. Electrospray ionisation parameters MS:MS_COMMENTS were: capillary +3 kV, sampling cone +40 V, source temperature 120 °C, MS:MS_COMMENTS desolvation temperature 550 °C, cone gas 100 L/h, desolvation gas 900 L/h, and MS:MS_COMMENTS nebulizer gas pressure 6.5 bar. Gas flows to the ion mobility region of the MS:MS_COMMENTS instrument were 2.0, 180 and 90 mL/min, for the trap, helium cell and IMS MS:MS_COMMENTS region, respectively. For the ozonolysis reactions, oxygen (≥ 99.999%, MS:MS_COMMENTS Coregas, NSW, Australia) was converted to ozone at concentrations of 220-240 MS:MS_COMMENTS g/Nm3 (~15-17% O3 in O2) using a high concentration ozone generator (TG-40, MS:MS_COMMENTS Ozone Solutions, Hull, IA). Peak areas for Criegee and aldehyde ions for each MS:MS_COMMENTS monounsaturated fatty acid isomer were combined and normalised to internal MS:MS_COMMENTS standard and total protein (measured by BCA assay). #MS_METABOLITE_DATA MS_METABOLITE_DATA:UNITS normalisd peak area MS_METABOLITE_DATA_START Samples M64_1 M64_2 M64_3 MIA_1 MIA_2 MIA_3 P150_1 P150_2 P150_3 PANC_1 PANC_2 PANC_3 Factors Sample source:MiaPaCa2 cells | group:GEMR Sample source:MiaPaCa2 cells | group:GEMR Sample source:MiaPaCa2 cells | group:GEMR Sample source:MiaPaCa2 cells | group:CON Sample source:MiaPaCa2 cells | group:CON Sample source:MiaPaCa2 cells | group:CON Sample source:Panc1 cells | group:GEMR Sample source:Panc1 cells | group:GEMR Sample source:Panc1 cells | group:GEMR Sample source:Panc1 cells | group:CON Sample source:Panc1 cells | group:CON Sample source:Panc1 cells | group:CON 16:1n-7 Precursor 4911440 4433769 4110076 5604789 7088610 7187151 3298023 6133222 6969477 5608628 4380095 6287812 16:1n-7 Criegee 103987 92644 86346 127752 163806 142836 55700 98087 125343 80083 60540 86765 16:1n-7 Aldehyde 188257 176388 159919 257358 325359 290576 107895 210154 245226 160837 124246 176238 16:1n-9 Precursor 4911440 4433769 4110076 5604789 7088610 7187151 3298023 6133222 6969477 5608628 4380095 6287812 16:1n-9 Criegee 28877 25847 27740 14108 16256 11401 14862 12769 15098 19397 13705 17178 16:1n-9 Aldehyde 45014 36210 42391 20367 28263 24914 28263 23371 29955 29918 22287 23158 16:1n-10 Precursor 4911440 4433769 4110076 5604789 7088610 7187151 3298023 6133222 6969477 5608628 4380095 6287812 16:1n-10 Criegee 3250 4731 4602 5005 5326 25808 10118 33569 30292 51441 40517 53449 16:1n-10 Aldehyde 7798 13323 9864 11858 13729 79371 27157 108741 81741 145769 116159 164050 18:1n-7 Precursor 19084702 18049386 17034088 17390728 18739616 19088110 15168191 17350246 20111674 20435610 17986006 21518846 18:1n-7 Criegee 424620 382325 325597 507677 576516 526628 437529 510085 657819 492482 388690 531993 18:1n-7 Aldehyde 714680 652173 539919 847588 981376 856802 654457 863582 1022688 770670 615659 846611 18:1n-9 Precursor 19084702 18049386 17034088 17390728 18739616 19088110 15168191 17350246 20111674 20435610 17986006 21518846 18:1n-9 Criegee 769771 720572 700932 555322 615759 603679 373498 452742 503228 563943 486648 634262 18:1n-9 Aldehyde 1475259 1373542 1337652 1063182 1188364 1161493 715192 895421 971775 1120753 959157 1245847 18:1n-10 Precursor 19084702 18049386 17034088 17390728 18739616 19088110 15168191 17350246 20111674 20435610 17986006 21518846 18:1n-10 Criegee 42204 41154 38135 33938 37257 60592 57453 133776 133828 221776 171641 237602 18:1n-10 Aldehyde 57021 60633 57406 58039 62192 111483 125147 283241 311193 501436 395453 541379 18:1n-12 Precursor 19084702 18049386 17034088 17390728 18739616 19088110 15168191 17350246 20111674 20435610 17986006 21518846 18:1n-12 Criegee 1576 1622 1591 1608 590 4747 2564 8451 3581 15126 10725 14277 18:1n-12 Aldehyde 7335 5796 6147 7290 7706 14699 4341 19962 11301 37613 35378 40973 19:0 Precursor 566186 523165 565939 469620 510180 713276 545260 666036 595391 519332 483600 624449 MS_METABOLITE_DATA_END #METABOLITES METABOLITES_START metabolite_name PrecursorMz ProductMz 16:1n-7 Precursor 421.3814 421.3814 16:1n-7 Criegee 421.3814 355.2679 16:1n-7 Aldehyde 421.3814 339.2787 16:1n-9 Precursor 421.3814 421.3814 16:1n-9 Criegee 421.3814 327.2402 16:1n-9 Aldehyde 421.3814 311.2473 16:1n-10 Precursor 421.3814 421.3814 16:1n-10 Criegee 421.3814 313.2293 16:1n-10 Aldehyde 421.3814 297.2331 18:1n-7 Precursor 449.4 449.4 18:1n-7 Criegee 449.4 383.2926 18:1n-7 Aldehyde 449.4 367.3047 18:1n-9 Precursor 449.4 449.4 18:1n-9 Criegee 449.4 355.2679 18:1n-9 Aldehyde 449.4 339.2787 18:1n-10 Precursor 449.4 449.4 18:1n-10 Criegee 449.4 341.2593 18:1n-10 Aldehyde 449.4 325.2615 18:1n-12 Precursor 449.4 449.4 18:1n-12 Criegee 449.4 313.2293 18:1n-12 Aldehyde 449.4 297.2331 19:0 Precursor 437.4068 437.4068 METABOLITES_END #END